2015 in paleontology

Paleontology or palaeontology (from Greek: paleo, "ancient"; ontos, "being"; and logos, "knowledge") is the study of prehistoric life forms on Earth through the examination of plant and animal fossils.[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2015.

Plants

Cnidarians

Name Novelty Status Authors Age Unit Location Notes Images

Acropora darrellae[2]

Sp. nov

Valid

Santodomingo, Wallace & Johnson

Miocene

 Indonesia

A staghorn coral, a species of Acropora.

Acropora elenae[2]

Sp. nov

Valid

Santodomingo, Wallace & Johnson

Miocene

 Indonesia

A staghorn coral, a species of Acropora.

Acropora emanuelae[2]

Sp. nov

Valid

Santodomingo, Wallace & Johnson

Miocene

 Indonesia

A staghorn coral, a species of Acropora.

Acropora hasibuani[2]

Sp. nov

Valid

Santodomingo, Wallace & Johnson

Miocene

 Indonesia

A staghorn coral, a species of Acropora.

Acropora laurae[2]

Sp. nov

Valid

Santodomingo, Wallace & Johnson

Late Oligocene

 Malaysia

A staghorn coral, a species of Acropora.

Acropora renemai[2]

Sp. nov

Valid

Santodomingo, Wallace & Johnson

Miocene

 Indonesia

A staghorn coral, a species of Acropora.

Actinaraea michoacanensis[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (early Aptian)

Cumburindio Formation

 Mexico

A stony coral belonging to the family Actinacididae, a species of Actinaraea.

Actinastrea chumbitaroensis[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Actinastraeidae, a species of Actinastrea.

Australastraea[4]

Gen. et comb. et 2 sp. nov

Valid

Denayer & Webb

Carboniferous (Mississippian)

 Australia

A rugose coral belonging to the group Stauriida and the family Lithostrotionidae. A new genus for "Lithostrotion" columnare Etheridge (1900); genus also contains "Orionastraea" columellaris Pickett (1966), "Lithostrotion" parvicolumnare Pickett (1966), "Lithostrotion" wilkinsoni Pickett (1966) and "Orionastraea" flemingi Webb (1990), as well as new species Australastraea arcifera and Australastraea carinata.

Axisvacuus tenerus[5]

Sp. nov

Valid

Fedorowski

Carboniferous (early Serpukhovian)

 Poland

A rugose coral belonging to the group Stauriida and the family Antiphyllidae, a species of Axisvacuus.

Birkenmajerites[5]

Gen. et sp. nov

Valid

Fedorowski

Carboniferous (early Serpukhovian)

 Poland

A rugose coral belonging to the group Stauriida and the family Ostravaiaidae. The type species is Birkenmajerites primus.

Cairnsipsammia[6]

Gen. et sp. nov

Valid

Baron-Szabo

Early Cretaceous (late Barremian–early Aptian)

Schrattenkalk Formation

 Austria

A stony coral belonging to the family Dendrophylliidae. The type species is C. merbeleri.

Chekhovichia[7]

Nom. nov

Valid

Doweld

Late Silurian

 Kyrgyzstan
 Russia
 Uzbekistan

An anthozoan belonging to the group Heliolitoidea and the family Innaporidae; a replacement name for Rotalites Leleshus (1970).

Chelmia[5]

Gen. et sp. nov

Valid

Fedorowski

Carboniferous (early Serpukhovian)

 Poland

A rugose coral belonging to the group Stauriida. The type species is Chelmia radiata.

Cionodendron smithi[4]

Sp. nov

Valid

Denayer & Webb

Carboniferous (early Viséan)

 Australia

A rugose coral belonging to the group Stauriida and the family Lithostrotionidae; a species of Cionodendron.

Craspedophyllia japonica[8]

Sp. nov

Valid

Stanley & Onoue

Late Triassic

 Japan

A stony coral belonging to the family Reimaniphylliidae, a species of Craspedophyllia.

Crassialveolites niigataensis[9]

Sp. nov

Valid

Niko, Ibaraki & Tazawa

Devonian

 Japan

A tabulate coral belonging to the family Alveolitidae.

Crateroseris stefani[10]

Sp. nov

Valid

EliĂĄĆĄovĂĄ

Late Jurassic (Tithonian)

Ć tramberk Limestone

 Czech Republic

A stony coral belonging to the family Synastraeidae, a species of Crateroseris.

Cryptosepta[11]

Gen. et sp. nov

Valid

Gretz, LathuiliĂšre & Martini

Early Jurassic (Hettangian)

 France

A zardinophyllid coral. The type species is Cryptosepta nuda.

Dendrophyllia okamotoi[12]

Sp. nov

Valid

Niko, Suzuki & Taguchi

Miocene

Katsuta Group

 Japan

A species of Dendrophyllia.

Echinophyllia sassellensis[13]

Sp. nov

Valid[14]

Budd & Bosellini

Oligocene

 Italy

Eocolumastrea[15]

Gen. et comb. nov

Valid

Löser & Zell

Cretaceous

 Romania

A stony coral belonging to the group Faviina and the family Columastraeidae. A new genus for "Columnocoenia" ksiazkiewiczi bucovinensis Morycowa (1971) (raised to the rank of a separate species Eocolumastrea bucovinensis)

Eokoninckocarinia gemmina[16]

Sp. nov

Valid

Denayer

Carboniferous (Mississippian)

Yaricak Formation

 Turkey

A rugose coral belonging to the group Stauriida, possibly a member of the family Koninckocariniidae.

Goniophyllum osloense[17]

Sp. nov

Valid

Johannessen in Wright

Silurian (late Telychian)

Vik Formation

 Norway

An operculate coral, a species of Goniophyllum.

Guembelastraea kanmerae[8]

Sp. nov

Valid

Stanley & Onoue

Late Triassic

 Japan

A stony coral belonging to the family Guembelastraeidae, a species of Guembelastraea.

Halucinophyllia[10]

Gen. et sp. nov

Valid

EliĂĄĆĄovĂĄ

Late Jurassic (Tithonian)

Ć tramberk Limestone

 Czech Republic

A stony coral belonging to the family Latomeandridae. The type species is Halucinophyllia subridens.

Heteropistophyllum carchensis[18]

Sp. nov

Valid

Löser, Arias & Vilas

Early Cretaceous (Aptian and Albian)

 Mexico
 Spain

A stony coral belonging to the family Elasmocoeniidae, a species of Heteropistophyllum.

Isopora matahari[2]

Sp. nov

Valid

Santodomingo, Wallace & Johnson

Miocene

 Indonesia

A staghorn coral, a species of Isopora.

Latiphyllia mexicana[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Montlivaltiidae, a species of Latiphyllia.

Leleshusia[19]

Nom. nov

Valid

Doweld

Late Ordovician

 China
 Kazakhstan
 Russia
 Tajikistan
 Uzbekistan

An anthozoan belonging to the group Heliolitoidea; a replacement name for Granulina Leleshus (1975).

Margarosmilia mizukamia[8]

Sp. nov

Valid

Stanley & Onoue

Late Triassic

 Japan

A stony coral belonging to the family Margarophylliidae, a species of Margarosmilia.

Mycetophyllopsis azteca[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Montlivaltiidae, a species of Mycetophyllopsis.

Munusculum[10]

Gen. et sp. nov

Valid

EliĂĄĆĄovĂĄ

Late Jurassic (Tithonian)

Ć tramberk Limestone

 Czech Republic

A stony coral belonging to the suborder Microsolenina. The type species is Munusculum martinaseki.

Nervophyllum lukoviensis[5]

Sp. nov

Valid

Fedorowski

Carboniferous (early Serpukhovian)

 Poland

A rugose coral belonging to the group Stauriida and the family Aulophyllidae, a species of Nervophyllum.

Nudacolumastrea[15]

Gen. et sp. nov

Valid

Löser & Zell

Cretaceous

 Spain

A stony coral belonging to the group Faviina and the family Columastraeidae. The type species is Nudacolumastrea stefani.

Occulogermen[5]

Gen. et sp. nov

Valid

Fedorowski

Carboniferous (early Serpukhovian)

 Poland

A rugose coral belonging to the group Stauriida and the family Lithostrotionidae. The type species is Occulogermen luciae.

Orbignygyra? incognita[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Dendrogyridae, possibly a species of Orbignygyra.

Ovalastrea malpaso[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Latomeandridae, a species of Ovalastrea.

Paraclausastrea vorarlbergensis[20]

Sp. nov

Valid

Baron-Szabo

Early Cretaceous (late Barremian-early Aptian)

Schrattenkalk Formation

 Austria

A montlivaltiid stony coral, a species of Paraclausastrea.

Paracycloseris effrenatus[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Cunnolitidae, a species of Paracycloseris.

Paraleptoria[13]

Gen. et comb. nov

Valid[14]

Budd & Bosellini

Eocene (Lutetian) to Oligocene (Rupelian)

 Italy

A new genus for "Meandrina" polygonalis Catullo (1856); genus also includes "Diploria" flexuosissima d'Achiardi (1868).

Parascolymia bracherti[21]

Sp. nov

Valid

Reuter in Reuter, Wiedl & Piller

Miocene (Langhian‒early Serravallian)

Leitha Limestone

 Austria

A stony coral belonging to the family Lobophylliidae, a species of Parascolymia.

Pickettodendron[4]

Gen. et comb. et sp. nov

Valid

Denayer & Webb

Carboniferous (Tournaisian)

 Australia

A rugose coral belonging to the group Stauriida and the family Lithostrotionidae. A new genus for "Lithostrotion" williamsi Pickett (1966); genus also contains "Schoenophyllum" dalmaensis Webb (1990), as well as new species Pickettodendron nudum.

Preverastraea coatlicueae[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Rhipidogyridae, a species of Preverastraea.

Preverastraea tociae[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Rhipidogyridae, a species of Preverastraea.

Pseudoroemeripora sugimurai[22]

Sp. nov

Valid

Niko, Fujikawa & Haikawa

Permian

Akiyoshi Limestone Group

 Japan

A tabulate coral.

Retiophyllia tosaensis[8]

Sp. nov

Valid

Stanley & Onoue

Late Triassic

 Japan

A stony coral belonging to the family Reimaniphylliidae, a species of Retiophyllia.

Rotiphyllum plumeum[5]

Sp. nov

Valid

Fedorowski

Carboniferous (early Serpukhovian)

 Poland

A rugose coral belonging to the group Stauriida and the family Antiphyllidae, a species of Rotiphyllum.

Sakalavastraea perturbata[23]

Sp. nov

Valid

Lösser & Sklenáƙ

Late Cretaceous (late Cenomanian)

Peruc-Korycany Formation

 Czech Republic

A stony coral belonging to the group Faviina.

Saltocyathus cumburindioensis[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (early Aptian)

Cumburindio Formation

 Mexico

A stony coral belonging to the family Rhipidogyridae, a species of Saltocyathus.

Sclerangia[24]

Gen. et sp. nov

Valid

Baron-Szabo & Cairns

Miocene

Chipola Formation

 United States

A rhizangiid stony coral. The type species is Sclerangia floridana.

?Sochkineophyllum symmetricum[5]

Sp. nov

Valid

Fedorowski

Carboniferous (early Serpukhovian)

 Poland

A rugose coral belonging to the group Stauriida and the family Polycoeliidae, possibly a species of Sochkineophyllum.

Sylviella[10]

Gen. et comb. et 4 sp. nov

Valid

EliĂĄĆĄovĂĄ

Late Jurassic (Tithonian)

Ć tramberk Limestone

 Czech Republic

A stony coral belonging to the suborder Caryophylliina. The type species is "Haplaraea" columnaris Ogilvie (1897); genus also contains new species Sylviella exquisita, Sylviella multisepta, Sylviella noveni and Sylviella benjamin.

Sylviellopsis[10]

Gen. et sp. nov

Valid

EliĂĄĆĄovĂĄ

Late Jurassic (Tithonian)

Ć tramberk Limestone

 Czech Republic

A stony coral belonging to the suborder Caryophylliina. The type species is Sylviellopsis erici.

Tegocoenia[10]

Gen. et comb. nov

Valid

EliĂĄĆĄovĂĄ

Late Jurassic (Tithonian)

Ć tramberk Limestone

 Czech Republic

A stony coral belonging to the family Placophylliidae. The type species is "Diplocoenia" jasenica FrajovĂĄ (1960).

Thalamocaeniopsis mexicanensis[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Thamnasteriidae, a species of Thalamocaeniopsis.

Thamnarea hornosensis[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (early Aptian)

Cumburindio Formation

 Mexico

A stony coral belonging to the family Actinacididae, a species of Thamnarea.

Thamnasteria tonantzinae[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Thamnasteriidae, a species of Thamnasteria.

Thamnasteriamorpha okudai[8]

Sp. nov

Valid

Stanley & Onoue

Late Triassic

 Japan

A stony coral belonging to the family Tropiastraeidae, a species of Thamnasteriamorpha.

Thecosmilia guerreroensis[3]

Sp. nov

Valid

Filkorn & Pantoja-Alor

Early Cretaceous (late Albian)

Mal Paso Formation

 Mexico

A stony coral belonging to the family Montlivaltiidae, a species of Thecosmilia.

Variabilifavia ausuganensis[13]

Sp. nov

Valid[14]

Budd & Bosellini

Oligocene (late Rupelian–early Chattian)

 Italy

Yakunopora mabutii[25]

Sp. nov

Valid

Niko

Permian (Wordian)

Iwaizaki Limestone

 Japan

A tabulate coral belonging to the group Favositida and the family Pachyporidae.

Zaphrentites rotiphylloides[5]

Sp. nov

Valid

Fedorowski

Carboniferous (early Serpukhovian)

 Poland

A rugose coral belonging to the group Stauriida and the family Stereophrentidae, a species of Zaphrentites.

Zaphriphyllum daleki[26]

Sp. nov

Valid

Denayer

Carboniferous (late Tournaisian)

Yılanlı Formation

 Turkey

A rugose coral belonging to the family Ekvasophyllidae, a species of Zaphriphyllum.

Zaphrufimia anceps[5]

Sp. nov

Valid

Fedorowski

Carboniferous (early Serpukhovian)

 Poland

A rugose coral belonging to the group Stauriida and the family Stereophrentidae, a species of Zaphrufimia.

Arthropods

Bryozoans

Name Novelty Status Authors Age Unit Location Notes Images

Acantholunaria[27]

Gen. et sp. nov

Valid

Ernst, Munnecke & Oswald

Silurian (early Wenlock)

 Sweden

A cystoporate bryozoan. The type species is Acantholunaria expansa.

Adeonellopsis incompta[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An adeonid cheilostome bryozoan, a species of Adeonellopsis.

Amphiblestrum paleogenicum[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A calloporid cheilostome bryozoan, a species of Amphiblestrum.

Amplexopora crassiparietum[30]

Sp. nov

Valid

Ernst & Nakrem

Silurian (Wenlock)

Steinsfjorden Formation

 Norway

An amplexoporid trepostome bryozoan, a species of Amplexopora.

Amplexopora evae[30]

Sp. nov

Valid

Ernst & Nakrem

Silurian (Wenlock)

Steinsfjorden Formation

 Norway

An amplexoporid trepostome bryozoan, a species of Amplexopora.

Andreella dubia[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A microporid cheilostome bryozoan, a species of Andreella.

Anaphragma undulata[31]

Sp. nov

Valid

Jiménez-Sånchez, Vennin & Villas

Ordovician (late Katian)

 Morocco

A trepostomate bryozoan, a species of Anaphragma.

Aostipora elongata[32]

Sp. nov

Valid

Jiménez-Sånchez in Jiménez-Sånchez, Villas & Vennin

Ordovician (late Katian)

Khabt-el-Hajar Formation

 Morocco

A trepostomate bryozoan, a species of Aostipora.

Arachnopusia dimorpha[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An arachnopusiid cheilostome bryozoan, a species of Arachnopusia.

Arthropoma renipora[33]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A lacernid schizoporelloid bryozoan, a species of Arthropoma.

Aspidostoma clava[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An aspidostomatid cheilostome bryozoan, a species of Aspidostoma.

Aspidostoma gelasinus[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An aspidostomatid cheilostome bryozoan, a species of Aspidostoma.

Aspidostoma twinn[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An aspidostomatid cheilostome bryozoan, a species of Aspidostoma.

Atactoporella moroccoensis[31]

Sp. nov

Valid

Jiménez-Sånchez, Vennin & Villas

Ordovician (late Katian)

 Morocco

A trepostomate bryozoan, a species of Atactoporella.

Baculopora redondensis[34]

Sp. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France

A member of Fenestrata belonging to the family Acanthocladiidae; a species of Baculopora.

Bryopesanser bragai[33]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian to Serravallian)

 Indonesia

An escharinid schizoporelloid bryozoan, a species of Bryopesanser.

Bryopesanser sanfilippoae[33]

Sp. nov

Valid

Di Martino & Taylor

Miocene (Burdigalian-Langhian boundary to Serravallian)

 Indonesia

An escharinid schizoporelloid bryozoan, a species of Bryopesanser.

Buffonellaria sagittaria[33]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A celleporid celleporoid bryozoan, a species of Buffonellaria.

Caberoides gordoni[33]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A catenicellid bryozoan, a species of Caberoides.

Cavernella asturica[35]

Sp. nov

Valid

Suårez Andrés & Ernst

Devonian

Moniello Formation

 Spain

A member of the family Fenestellidae.

Celiopsis[36]

Gen. et sp. et comb. nov

Valid

ZĂĄgorĆĄek, Gordon & VĂĄvra

Eocene to Oligocene

 Austria
 Hungary
 Italy
 Poland
 Slovakia
 Spain

A chlidoniopsid cheilostome bryozoan. The type species is Celiopsis vici; genus also contains "Crisidia" vindobonensis Reuss (1847) and "Chlidoniopsis" vavrai ZĂĄgorĆĄek (2003).

Cellaria bicuspidata[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A cellariid cheilostome bryozoan, a species of Cellaria.

Cellaria gigas[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A cellariid cheilostome bryozoan, a species of Cellaria.

Cellaria inarticulata[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A cellariid cheilostome bryozoan, a species of Cellaria.

Cellaria palatum[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A cellariid cheilostome bryozoan, a species of Cellaria.

Chaperiopsis cookae[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A chaperiid cheilostome bryozoan, a species of Chaperiopsis.

Chataimulosia revelator[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A lepraliellid cheilostome bryozoan, a species of Chataimulosia.

Cladobryopastor[28]

Gen. et sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A bryopastorid cheilostome bryozoan. The type species is Cladobryopastor philipi.

Conopeum stamenocelloides[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A cheilostome bryozoan. Originally described as a species of Conopeum; Dick, Sakamoto & Komatsu (2018) transferred this species to the genus Kenocharixa.[37]

Curvipora[27]

Gen. et sp. nov

Valid

Ernst, Munnecke & Oswald

Silurian (early Wenlock)

 Sweden

A cystoporate bryozoan. The type species is Curvipora monostylata.

Cyphotrypa regularis[32]

Sp. nov

Valid

Jiménez-Sånchez in Jiménez-Sånchez, Villas & Vennin

Ordovician (late Katian)

Khabt-el-Hajar Formation

 Morocco

A trepostomate bryozoan, a species of Cyphotrypa.

Cystodictya gallensis[34]

Sp. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France

A cystoporate bryozoan belonging to the family Cystodictyonidae; a species of Cystodictya.

Dactylostega spiralis[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A foveolariid cheilostome bryozoan, a species of Dactylostega.

Diedropora[28]

Gen. et sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An adeonoid cheilostome bryozoan. The type species is Diedropora gravabilis.

Dybowskiella piriforme[34]

Sp. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France

A cystoporate bryozoan belonging to the family Fistuliporidae; a species of Dybowskiella.

Dybowskiella rotunda[34]

Sp. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France

A cystoporate bryozoan belonging to the family Fistuliporidae; a species of Dybowskiella.

Dyscritella ornata[38]

Sp. nov

Valid

Tolokonnikova, Ernst & Poty

Carboniferous (Tournaisian)

Namur-Dinant Basin

 Belgium

Dyscritella perforata[39]

Sp. nov

Valid

Tolokonnikova et al.

Devonian (latest Famennian)

Comblain-au-Pont Formation

 Belgium

A dyscritellid trepostome bryozoan, a species of Dyscritella.

Eridopora suarezi[34]

Sp. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France

A cystoporate bryozoan belonging to the family Fistuliporidae; a species of Eridopora.

Exechonella chathamensis[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An exechonellid cheilostome bryozoan, a species of Exechonella.

Exochella abigailae[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A romancheinid cheilostome bryozoan, a species of Exochella.

Exochella linearis[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A romancheinid cheilostome bryozoan, a species of Exochella.

Exochella reidae[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A romancheinid cheilostome bryozoan, a species of Exochella.

Exochella woodae[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A romancheinid cheilostome bryozoan, a species of Exochella.

Fabifenestella macrofenestrata[34]

Sp. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France

A member of Fenestrata belonging to the family Fenestellidae; a species of Fabifenestella.

?Filaguria kalimantanensis[33]

Sp. nov

Valid

Di Martino & Taylor

Miocene (Serravallian)

 Indonesia

A cribrilinid bryozoan, possibly a species of Filaguria.

Fistulipora tolokonnikovae[34]

Sp. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France

A cystoporate bryozoan belonging to the family Fistuliporidae; a species of Fistulipora.

Floridina elegans[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An onychocellid cheilostome bryozoan, a species of Floridina.

Gephyrotes moissettei[40]

Sp. nov

Valid

Di Martino & Rosso

Miocene (Tortonian)

 Italy

A member of Ascophora belonging to the family Cribrilinidae.

Gigantopora milenae[33]

Sp. nov

Valid

Di Martino & Taylor

Miocene (Serravallian)

 Indonesia

A gigantoporid schizoporelloid bryozoan, a species of Gigantopora.

Gigantopora grandis[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A gigantoporid cheilostome bryozoan, a species of Gigantopora.

Gigantopora modesta[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A gigantoporid cheilostome bryozoan, a species of Gigantopora.

Gorjunopora[34]

Gen. et sp. et comb. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France
 Russia

A member of Fenestrata belonging to the family Acanthocladiidae. The type species is Gorjunopora gallica; genus also includes Gorjunopora triangulata (Schulga-Nesterenko, 1955).

Hemicyclopora dissidens[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A romancheinid cheilostome bryozoan, a species of Hemicyclopora.

Hemicyclopora ventricosa[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A romancheinid cheilostome bryozoan, a species of Hemicyclopora.

Heminematopora gaetula[41]

Sp. nov

Valid

Ernst, Jiménez-Sånchez & Baidder

Ordovician (Katian)

 Morocco

A species of Heminematopora.

Hemitrypa adversa[35]

Sp. nov

Valid

Suårez Andrés & Ernst

Devonian

Moniello Formation

 Spain

A member of the family Fenestellidae.

Hemitrypa terentei[35]

Sp. nov

Valid

Suårez Andrés & Ernst

Devonian

Moniello Formation

 Spain

A member of the family Fenestellidae.

Heteropora scholzi[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A cerioporid cyclostome bryozoan, a species of Heteropora.

Heterotrypa ringerikense[30]

Sp. nov

Valid

Ernst & Nakrem

Silurian (Wenlock)

Steinsfjorden Formation

 Norway

A heterotrypid trepostome bryozoan, a species of Heterotrypa.

?Hippomenella devatasae[33]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A romancheinid lepralielloid bryozoan, possibly a species of Hippomenella.

?Hippomenella uniserialis[33]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A romancheinid lepralielloid bryozoan, possibly a species of Hippomenella.

Hippopleurifera barbosae[42]

Sp. nov

Valid

Ramalho et al.

Miocene

Pirabas Formation

 Brazil

A cheilostome bryozoan belonging to the superfamily Lepralielloidea and the family Romancheneidae, a species of Hippopleurifera.

Hippopleurifera confusa[42]

Sp. nov

Valid

Ramalho et al.

Miocene

Pirabas Formation

 Brazil

A cheilostome bryozoan belonging to the superfamily Lepralielloidea and the family Romancheneidae, a species of Hippopleurifera.

Hippopodina indicata[33]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian to Langhian)

 Indonesia

A hippopodinid schizoporelloid bryozoan, a species of Hippopodina.

Hyporosopora nana[43]

Sp. nov

Valid

Wilson, Bosch & Taylor

Middle Jurassic (Callovian)

Matmor Formation

 Israel

A plagioeciid cyclostome bryozoan, a species of Hyporosopora.

Hyporosopora negevensis[43]

Sp. nov

Valid

Wilson, Bosch & Taylor

Middle Jurassic (Callovian)

Matmor Formation

 Israel

A plagioeciid cyclostome bryozoan, a species of Hyporosopora.

Idmonea snehi[43]

Sp. nov

Valid

Wilson, Bosch & Taylor

Middle Jurassic (Callovian)

Matmor Formation

 Israel

A multisparsid cyclostome bryozoan, a species of Idmonea.

Illusiopora[28]

Gen. et 2 sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A schizoporelloid cheilostome bryozoan. The type species is Illusiopora bifax; genus also contains Illusiopora recta.

Isostylus veserensis[44]

Sp. nov

Valid

Ernst, Tolokonnikova & Denayer

Devonian (late Frasnian)

 Belgium

A species of Isostylus.

Lacerna ordinaria[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A lacernid cheilostome bryozoan, a species of Lacerna.

Lagenipora sciutoi[33]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A celleporid celleporoid bryozoan, a species of Lagenipora.

Lenapora gurievensis[45]

Sp. nov

Valid

Mesentseva

Devonian

 Russia

A rhabdomesid bryozoan, a species of Lenapora.

Leptotrypa enodis[27]

Sp. nov

Valid

Ernst, Munnecke & Oswald

Silurian (early Wenlock)

 Sweden

A trepostome bryozoan, a species of Leptotrypa.

Leptotrypa hexagona[38]

Sp. nov

Valid

Tolokonnikova, Ernst & Poty

Carboniferous (Tournaisian)

Namur-Dinant Basin

 Belgium

Leptotrypa perforata[27]

Sp. nov

Valid

Ernst, Munnecke & Oswald

Silurian (early Wenlock)

 Sweden

A trepostome bryozoan, a species of Leptotrypa.

Macrocamera obesa[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An eminooeciid cheilostome bryozoan, a species of Macrocamera.

Margaretta amitabhae[33]

Sp. nov

Valid

Di Martino & Taylor

Miocene (Burdigalian-Langhian boundary to Serravallian)

 Indonesia

A margarettid schizoporelloid bryozoan, a species of Margaretta.

Marssonopora connexa[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A calloporid cheilostome bryozoan, a species of Marssonopora.

Mediaporina kiaeri[30]

Sp. nov

Valid

Ernst & Nakrem

Silurian (Wenlock)

Steinsfjorden Formation

 Norway

An arthrostylid cryptostome bryozoan, a species of Mediaporina.

Megacanthopora enodata[34]

Sp. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France

A member of Cryptostomata belonging to the family Rhomboporidae; a species of Megacanthopora.

Melychocella bilamellata[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An aspidostomatid cheilostome bryozoan, a species of Melychocella.

Melychocella obliqua[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An aspidostomatid cheilostome bryozoan, a species of Melychocella.

Mesenteripora jamesi[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A plagioeciid cyclostome bryozoan, a species of Mesenteripora.

Microeciella yoavi[43]

Sp. nov

Valid

Wilson, Bosch & Taylor

Middle Jurassic (Callovian)

Matmor Formation

 Israel

An oncousoeciid cyclostome bryozoan, a species of Microeciella.

Micropora chathamica[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A microporid cheilostome bryozoan, a species of Micropora.

Monticulipora globulata[31]

Sp. nov

Valid

Jiménez-Sånchez, Vennin & Villas

Ordovician (late Katian)

 Morocco

A trepostomate bryozoan, a species of Monticulipora.

Monticulipora irregularis[31]

Sp. nov

Valid

Jiménez-Sånchez, Vennin & Villas

Ordovician (late Katian)

 Morocco

A trepostomate bryozoan, a species of Monticulipora.

Moyanopora[28]

Gen. et sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An arachnopusiid cheilostome bryozoan. The type species is Moyanopora hugoi.

Multescharellina pisiformis[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A lepraliellid cheilostome bryozoan, a species of Multescharellina.

Myriapora sciutoi[46]

Sp. nov

Valid

Rosso & Di Martino

Miocene (Tortonian and Messinian) to Pleistocene (Gelasian)

 Italy
 Morocco

A member of Neocheilostomina belonging to the family Myriaporidae; a species of Myriapora.

Nematopora rara[45]

Sp. nov

Valid

Mesentseva

Devonian

 Russia

A rhabdomesid bryozoan, a species of Nematopora.

Nikiforovastylus[39]

Gen. et sp. nov

Valid

Tolokonnikova et al.

Devonian (latest Famennian)

Comblain-au-Pont Formation

 Belgium

A nikiforovellid cryptostome bryozoan. The type species is Nikiforovastylus ourthensis.

Nikiforovella dinantia[39]

Sp. nov

Valid

Tolokonnikova et al.

Devonian (latest Famennian)

Comblain-au-Pont Formation

 Belgium

A nikiforovellid cryptostome bryozoan, a species of Nikiforovella.

Nikiforovella tobolensis[47]

Sp. nov

Valid

Tolokonnikova

Carboniferous (Tournaisian)

 Russia

Ogiva ellinorvoigtae[48]

Sp. nov

Valid[49]

Martha et al.

Cretaceous (middle Santonian)

Gehrden Formation

 Germany

An onychocellid cheilostome bryozoan, a species of Ogiva.

Onychocella exilis[50]

Sp. nov

Valid

Koromyslova & Shcherbinina

Late Cretaceous (Maastrichtian)

 Uzbekistan

An onychocellid cheilostome bryozoan, a species of Onychocella.

Onychocella subtriangulata[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An onychocellid cheilostome bryozoan, a species of Onychocella.

Orthopora worsleyi[30]

Sp. nov

Valid

Ernst & Nakrem

Silurian (Wenlock)

Steinsfjorden Formation

 Norway

A rhabdomesid cryptostome bryozoan, a species of Orthopora.

Osthimosia aurora[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A celleporid cheilostome bryozoan, a species of Osthimosia.

Osthimosia curiosa[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A celleporid cheilostome bryozoan, a species of Osthimosia.

Oviexechonella[33]

Gen. et sp. nov

Valid

Di Martino & Taylor

Miocene (Serravallian)

 Indonesia

An exechonellid arachnopusioid bryozoan. The type species is Oviexechonella digeronimoi.

Paracuneatopora egregia[45]

Sp. nov

Valid

Mesentseva

Devonian

 Russia

A rhabdomesid bryozoan, a species of Paracuneatopora.

Paracuneatopora kuvaschensis[45]

Sp. nov

Valid

Mesentseva

Devonian

 Russia

A rhabdomesid bryozoan, a species of Paracuneatopora.

Paranicklesopora ornaisa[38]

Sp. nov

Valid

Tolokonnikova, Ernst & Poty

Carboniferous (Tournaisian)

Namur-Dinant Basin

 Belgium

Parvohallopora cystata[32]

Sp. nov

Valid

Jiménez-Sånchez in Jiménez-Sånchez, Villas & Vennin

Ordovician (late Katian)

Khabt-el-Hajar Formation

 Morocco

A trepostomate bryozoan, a species of Parvohallopora.

Perkhurovella[51]

Gen. et sp. nov

Valid

Gorjunova

Carboniferous

 Russia

A diploporariine acanthocladiid bryozoan. The type species is Perkhurovella clara.

Phonicosia sinuosa[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A lacernid cheilostome bryozoan, a species of Phonicosia.

Plagioecia zatoni[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A plagioeciid cyclostome bryozoan, a species of Plagioecia.

Platonea dilatata[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A tubuliporid cyclostome bryozoan, a species of Platonea.

Porella tiorioriensis[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A bryocryptellid cheilostome bryozoan, a species of Porella.

Porina turrita[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A porinid cheilostome bryozoan, a species of Porina.

Primorella variata[47]

Sp. nov

Valid

Tolokonnikova

Carboniferous (Tournaisian)

 Russia

Prophyllodictya simplex[52]

Sp. nov

Valid[53]

Ma et al.

Ordovician (early Tremadocian)

Nantzinkuan Formation

 China

A cryptostome bryozoan, a species of Prophyllodictya.

Pseudobatostomella parva[39]

Sp. nov

Valid

Tolokonnikova et al.

Devonian (latest Famennian)

Comblain-au-Pont Formation

 Belgium

A dyscritellid trepostome bryozoan, a species of Pseudobatostomella.

Pseudothyracella campbelli[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A bryopastorid cheilostome bryozoan, a species of Pseudothyracella.

Puellina bontangensis[33]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A cribrilinid bryozoan, a species of Puellina.

Pyrisinella primazelandiae[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A pyrisinellid cheilostome bryozoan, a species of Pyrisinella.

Quasitrilaminopora[28]

Gen. et sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

An arachnopusiid cheilostome bryozoan. The type species is Quasitrilaminopora curiosa.

Radiotrypa alnifensis[41]

Sp. nov

Valid

Ernst, Jiménez-Sånchez & Baidder

Ordovician (Katian)

 Morocco

A species of Radiotrypa.

Rectifenestella covae[35]

Sp. nov

Valid

Suårez Andrés & Ernst

Devonian

Moniello Formation

 Spain

A member of the family Fenestellidae.

Reptadeonella curvabilis[33]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

An adeonid bryozoan, a species of Reptadeonella.

Reptadeonella toddi[33]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian or early Langhian to Serravallian)

 Indonesia

An adeonid bryozoan, a species of Reptadeonella.

Reteporella mediocris[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A phidoloporid cheilostome bryozoan, a species of Reteporella.

Revalotrypa ramosa[41]

Sp. nov

Valid

Ernst, Jiménez-Sånchez & Baidder

Ordovician (Katian)

 Morocco

A species of Revalotrypa.

Saevitella renemai[33]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A hippopodinid schizoporelloid bryozoan, a species of Saevitella.

Saffordotaxis spinigerus[38]

Sp. nov

Valid

Tolokonnikova, Ernst & Poty

Carboniferous (Tournaisian)

Namur-Dinant Basin

 Belgium

Salairella[45]

Gen. et 5 sp. nov

Junior homonym

Mesentseva

Devonian

 Russia

A rhabdomesid bryozoan. Genus contains five species: S. petaloida, S. variabilis, S. maculata, S. baskuskanensis and S. recta. The generic name is a junior homonym of Salairella Khalfina (1961) and Salairella Severgina (1984).

Schizoretepora tamagawensis[54]

Sp. nov

Valid

ZĂĄgorĆĄek, Takashima & Hirose

Miocene

Tanosawa Formation

 Japan

A member of Phidoloporidae, a species of Schizoretepora.

Sendinopora[33]

Gen. et sp. nov

Valid

Di Martino & Taylor

Miocene (Serravallian)

 Indonesia

A tessaradomid lepralielloid bryozoan. The type species is Sendinopora prima.

Siphonicytara litotes[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A siphonicytarid cheilostome bryozoan, a species of Siphonicytara.

Siphonicytara primitiva[28]

Sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A siphonicytarid cheilostome bryozoan, a species of Siphonicytara.

Spinofenestella nodosa[38]

Sp. nov

Valid

Tolokonnikova, Ernst & Poty

Carboniferous (Tournaisian)

Namur-Dinant Basin

 Belgium

Stellatotrypa[27]

Gen. et sp. nov

Valid

Ernst, Munnecke & Oswald

Silurian (early Wenlock)

 Sweden

A trepostome bryozoan. The type species is Stellatotrypa hirsuta.

Sulcoretepora regularis[47]

Sp. nov

Valid

Tolokonnikova

Carboniferous (Tournaisian)

 Russia

Terepennipora[51]

Gen. et comb. nov

Valid

Gorjunova

Carboniferous

 Russia

A diploporariine acanthocladiid bryozoan. A new genus for "Pinnatopora" longicellata Morozova (1955); genus also contains "Pinnatopora" volgensis Shishova (1959), "Pinnatopora" inconstans Shishova (1959), "Pinnatopora" subpunctata Shishova (1959) and "Pinnatopora" distincta Shishova (1950).

Trematopora maculata[30]

Sp. nov

Valid

Ernst & Nakrem

Silurian (Wenlock)

Steinsfjorden Formation

 Norway

A trematoporid trepostome bryozoan, a species of Trematopora.

Triznotrypa potii[44]

Sp. nov

Valid

Ernst, Tolokonnikova & Denayer

Devonian (late Frasnian)

 Belgium

A species of Triznotrypa.

Trypostega hasibuani[33]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A trypostegid hippothooid bryozoan, a species of Trypostega.

Tubiporella magnipora[33]

Sp. nov

Valid

Di Martino & Taylor

Miocene (late Burdigalian)

 Indonesia

A didymosellid didymoselloid bryozoan, a species of Tubiporella.

Tumaiella[28]

Gen. et sp. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 New Zealand

A calloporid cheilostome bryozoan. The type species is Tumaiella dieffenbachi.

Volgia deftera[34]

Sp. nov

Valid

Ernst, Wyse Jackson & Aretz

Carboniferous (Viséan)

 France

A cystoporate bryozoan belonging to the family Hexagonellidae; a species of Volgia.

Zagorsekia[28]

Gen. et comb. nov

Valid[29]

Gordon & Taylor

Early Eocene

Tumaio Limestone

 Australia
 New Zealand

A tubuliporid cyclostome bryozoan; a new genus for "Heteropora" nodulosa MacGillivray (1895).

Brachiopods

Name Novelty Status Authors Age Unit Location Notes Images

Alaskothyris[55]

Gen. et sp. nov

Valid

Blodgett, Baranov & Santucci

Devonian (Givetian)

 United States

A stringocephalid terebratulid brachiopod. The type species is Alaskothyris frosti.

Albasphe[56]

Gen. et sp. nov

Valid

Halamski et al.

Middle Triassic (Ladinian)

 Croatia

A member of Terebratulida. The type species is Albasphe albertimagni.

Alexenia? delepinei[57]

Sp. nov

Valid

ChacĂłn & Winkler Prins

Carboniferous (Pennsylvanian)

 Spain

A member of Productidae, possibly a species of Alexenia.

Amictocracens? brocki[58]

Sp. nov

Valid

Popov et al.

Cambrian

Parahio Formation

 India

A member of Acrotretida belonging to the family Acrotretidae, possibly a species of Amictocracens.

Aphelotreta khemangarensis[58]

Sp. nov

Valid

Popov et al.

Cambrian

Parahio Formation

 India

A member of Acrotretida belonging to the family Acrotretidae, a species of Aphelotreta.

Auriculispina kanmerai[59]

Sp. nov

Valid

Tazawa & Shintani

Permian (Sakmarian)

Sakamotozawa Formation

 Japan

A member of Productida belonging to the family Kansuellidae.

Bifida rara[60]

Sp. nov

Valid

Baranov

Devonian (Emsian)

 Russia

A member of Atrypidae, a species of Bifida.

Bockelia canalis[61]

Sp. nov

Valid

Benedetto

Ordovician (Darriwilian)

Los Azules Formation

 Argentina

A member of Strophomenida belonging to the superfamily Plectambonitoidea and the family Taffidae; a species of Bockelia.

Castrillonia[62]

Gen. et sp. nov

Valid

GarcĂ­a-Alcalde

Devonian

 Spain

A member of Orthida belonging to the family Rhipidomellidae. The type species is C. vivaldiana.

Chilcorthis[61]

Gen. et sp. nov

Valid

Benedetto

Ordovician (Darriwilian)

Los Azules Formation

 Argentina

A member of Orthida belonging to the superfamily Plectorthoidea. The type species is Chilcorthis huarpe.

Chonetinetes elongatus[63]

Sp. nov

Valid

Afanasjeva, Jun-ichi & Tomohiko

Permian

 Japan

A member of Chonetida belonging to the family Rugosochonetidae, a species of Chonetinetes.

Chopinia[64]

Gen. et sp. nov

Junior homonym

GarcĂ­a-Alcalde & Herrera

Devonian (late Lochkovian or the early Pragian)

 Spain

A member of Athyridida belonging to the family Athyrididae. The type species is C. petercarlsi. The generic name is preoccupied by Chopinia d'Abrera (2001).

Chynithele intermedia[65]

Sp. nov

Valid

Mergl & Jiménez-Sånchez

Devonian (late Emsian)

Daleje-Tƙebotov Formation

 Czech Republic

A member of Discinidae, a species of Chynithele.

Copperella[60]

Gen. et sp. nov

Valid

Baranov

Devonian (Emsian)

 Russia

A member of Atrypidae. The type species is Copperella kaskevichi.

Cyrtonotreta vasculata[61]

Sp. nov

Valid

Benedetto

Ordovician (Darriwilian)

Los Azules Formation

 Argentina

A member of Acrotretida belonging to the family Acrotretidae; a species of Cyrtonotreta.

Dagyssia[66]

Gen. et comb. nov

Valid

Gaetani & Mantovani

Triassic (Anisian)

 Croatia
 Hungary
 Italy
 China?

A mentzeliine mentzeliid brachiopod. The type species is "Spiriferina" palaeotypus Loretz (1875); genus might also contain "Mentzelia" multicostata Yang & Xu (1966).

Dalejina telemanni[62]

Sp. nov

Valid

GarcĂ­a-Alcalde

Devonian

 Spain

A member of Orthida belonging to the family Rhipidomellidae.

Drahanorhynchus gleniffensis[67]

Sp. nov

Valid

Mottequin, Sevastopulo & Simon

Carboniferous (Viséan)

Meenymore Formation

 Ireland

A member of Orthotetida belonging to the family Schuchertellidae.

Duartea latoresensis[57]

Sp. nov

Valid

ChacĂłn & Winkler Prins

Carboniferous (Pennsylvanian)

 Spain

A member of Productidae, possibly a species of Duartea.

Eodmitria sezgini[68]

Sp. nov

Valid

Gourvennec & HoƟgör

Devonian (latest Givetian to early Frasnian)

GĂŒmĂŒÈ™ali Formation

 Turkey

A member of Spiriferida belonging to the family Cyrtospiriferidae, a species of Eodmitria.

Eohadrotreta haydeni[58]

Sp. nov

Valid

Popov et al.

Cambrian

Parahio Formation

 India

A member of Acrotretida belonging to the family Acrotretidae, a species of Eohadrotreta.

Eucharitina bultyncki[64]

Sp. nov

Valid

GarcĂ­a-Alcalde & Herrera

Devonian (early Lochkovian)

 Spain

A member of Rhynchonellida belonging to the family Uncinulidae.

Eucharitina carlsi[64]

Sp. nov

Valid

GarcĂ­a-Alcalde & Herrera

Devonian (early Pragian)

 Spain

A member of Rhynchonellida belonging to the family Uncinulidae.

Eurytreta harringtoni[69]

Sp. nov

Valid

Mergl & Herrera in Mergl et al.

Late Cambrian

Lampazar Formation

 Argentina

Evanidisinurostrum saouraense[70]

Sp. nov

Valid

Mottequin et al.

Devonian (Famennian)

 Algeria

A rhynchonellid brachiopod, a species of Evanidisinurostrum.

Hadrotreta timchristiorum[58]

Sp. nov

Valid

Popov et al.

Cambrian

Parahio Formation

 India

A member of Acrotretida belonging to the family Acrotretidae, a species of Hadrotreta.

Hibernodonta lakhensis[71]

Sp. nov

Valid

Popov, Holmer & Baars in Popov et al.

Ordovician (Katian)

Chahgonbad Formation

 Iran

A rafinesquinid strophomenid brachiopod; a species of Hibernodonta.

Hindella prima[71]

Sp. nov

Valid

Popov, Holmer & Baars in Popov et al.

Ordovician (Katian)

Chahgonbad Formation

 Iran

A meristellid athyridid brachiopod; a species of Hindella.

Inflatia ovetensis[57]

Sp. nov

Valid

ChacĂłn & Winkler Prins

Carboniferous (Pennsylvanian)

 Spain

A member of Productidae, possibly a species of Inflatia.

Jafarirhynchus[72]

Gen. et sp. nov

Valid

Popov et al.

Silurian (Telychian)

Niur Formation

 Iran

A rhynchotrematoid rhynchonellid brachiopod. The type species is Jafarirhynchus alatus.

Kerberellus[73]

Gen. et sp. nov

Valid

Devaere, Holmer & Clausen in Devaere et al.

Cambrian (Terreneuvian)

Marcou Formation

 France

A mickwitziid-like stem-group brachiopod. The type species is Kerberellus marcouensis.

Kirkirhynchus[74]

Gen. et comb. nov

Valid

Baranov & Blodgett

Devonian (early Pragian)

 United States

A trigonirhynchiid rhynchonellid brachiopod; a new genus for "Camarotoechia" reesidei Kirk & Amsden (1952).

Kosagittella robusta[65]

Sp. nov

Valid

Mergl & Jiménez-Sånchez

Devonian (late Emsian)

Daleje-Tƙebotov Formation

 Czech Republic

A member of Obolidae, a species of Kosagittella.

Ktenopotamorhynchus[75]

Gen. et comb. nov

Valid

Sartenaer

Devonian (Lochkovian)

 Czech Republic
 Ukraine

A member of Rhynchonellida. The type species is "Rhynchonella" borealiformis Siemiradzki (1906); genus also includes "Plethorhyncha" altera (Barrande, 1879).

Kutorgina songshanensis[76]

Sp. nov

Valid

Liu et al.

Cambrian

 China

A species of Kutorgina.

Leptembolon argentinum[77]

Sp. nov

Valid

Benedetto & Muñoz

Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A member of Obolidae, a species of Leptembolon.

Libecoviella lenticularis[69]

Sp. nov

Valid

Mergl & Herrera in Mergl et al.

Late Cambrian

Lampazar Formation

 Argentina

Libecoviella tilcarensis[77]

Sp. nov

Valid

Benedetto & Muñoz

Ordovician (Tremadocian)

Santa Rosita Formation

 Argentina

A member of Obolidae, a species of Libecoviella.

Lingularia salymica[78]

Sp. nov

Valid

Smirnova in Smirnova et al.

Late Jurassic

 Russia

A member of Lingulida, a species of Lingularia.

Lingulella? melonica[69]

Sp. nov

Valid

Mergl & Herrera in Mergl et al.

Late Cambrian

Lampazar Formation

 Argentina

Lochkothele rugellata[65]

Sp. nov

Valid

Mergl & Jiménez-Sånchez

Devonian (late Emsian)

Daleje-Tƙebotov Formation
ZlĂ­chov Formation

 Czech Republic

A member of Discinidae, a species of Lochkothele.

Lomatorthis? multilamellosa[71]

Sp. nov

Valid

Popov, Holmer & Baars in Popov et al.

Ordovician (Darriwilian)

Chahgonbad Formation

 Iran

A glyptorthid orthid brachiopod; possibly a species of Lomatorthis.

Maisorthina[62]

Gen. et sp. nov

Valid

GarcĂ­a-Alcalde

Devonian (late Emsian)

 Spain

A member of Orthida belonging to the family Platyorthidae. The type species is M. saroyani.

Mariyaspiriferella[79]

Gen. et sp. et comb. nov

Valid

Baranov & Blodgett

Early Devonian

 Canada
 Russia
 United States

A cyrtinid brachiopod. The type species is M. sibirica; genus also includes "Cyrtina" impressio Perry (1984).

Misensia[60]

Gen. et sp. nov

Valid

Baranov

Devonian (Emsian)

 Russia

A member of Atrypidae. The type species is Misensia gracilis.

Neochonetes (Zhongyingia) transversa[80]

Sp. nov

Valid

Zhang et al.

Permian (Lopingian)

Longtan Formation
Yanshi Formation

 China

A member of Productida belonging to the family Rugosochonetidae.

Ogilviella gerensis[60]

Sp. nov

Valid

Baranov

Devonian (Emsian)

 Russia

A member of Atrypidae, a species of Ogilviella.

Paillettia menae[68]

Sp. nov

Valid

Gourvennec & HoƟgör

Devonian (latest Givetian to early Frasnian)

GĂŒmĂŒÈ™ali Formation

 Turkey

A member of Spiriferida belonging to the family Adolfiidae, a species of Paillettia.

Palaeoglossa minima[61]

Sp. nov

Valid

Benedetto

Ordovician (Darriwilian)

Los Azules Formation

 Argentina

A member of Obolidae; a species of Palaeoglossa.

Parmorthina gozonica[62]

Sp. nov

Valid

GarcĂ­a-Alcalde

Devonian

 Spain

A member of Orthida belonging to the family Dalmanellidae.

Paryphella acutula[80]

Sp. nov

Valid

Zhang et al.

Permian (Lopingian)

Dalong Formation
Longtan Formation

 China

A member of Productida belonging to the family Productellidae.

Pelmanotreta[81]

Nom. nov

Valid

Skovsted et al.

Early Cambrian

 Russia

A replacement name for Cryptotreta Pelman (1977) (preoccupied).

Philhedra pauciradiata[61]

Sp. nov

Valid

Benedetto

Ordovician (Darriwilian)

Los Azules Formation

 Argentina

A member of Craniidae; a species of Philhedra.

Portrania[82]

Gen. et sp. nov

Valid[83]

Cocks

Ordovician (late Katian)

Portrane Limestone

 Ireland

A chilidiopsoidean brachiopod. The type species is Portrania wrighti Cocks & Harper in Cocks (2015).

Praeoehlertella lukesi[65]

Sp. nov

Valid

Mergl & Jiménez-Sånchez

Devonian (late Emsian)

Daleje-Tƙebotov Formation

 Czech Republic

A member of Discinidae, a species of Praeoehlertella.

Prototreta? sumnaensis[58]

Sp. nov

Valid

Popov et al.

Cambrian

Parahio Formation

 India

A member of Acrotretida belonging to the family Acrotretidae, possibly a species of Prototreta.

Psilothyris corbachoi[84]

Sp. nov

Valid

Calzada & Moreno

Early Cretaceous (Aptian)

 Spain

A member of Terebratellidina belonging to the family Laqueidae and the subfamily Terebrataliopsinae, a species of Psilothyris.

Ptychomentzelia[66]

Gen. et comb. et sp. nov

Valid

Gaetani & Mantovani

Middle Triassic

 Austria
 Bosnia and Herzegovina
 Hungary
 Italy
 Romania
 Turkey

A tethyspirine mentzeliid brachiopod. The type species is "Spiriferina" propontica Toula (1896); genus also contains "Spiriferina" ptychitiphila Bittner (1890), as well as the new species Ptychomentzelia ritensis.

Raisiya[60]

Gen. et sp. nov

Valid

Baranov

Devonian (Emsian)

 Russia

A member of Atrypidae. The type species is Raisiya solonchanensis.

Rhipidomella cervantesi[62]

Sp. nov

Valid

GarcĂ­a-Alcalde

Devonian

 Spain

A member of Orthida belonging to the family Rhipidomellidae.

Rugaria semicircularis[63]

Sp. nov

Valid

Afanasjeva, Jun-ichi & Tomohiko

Permian

 Japan

A member of Chonetida belonging to the family Rugosochonetidae, a species of Rugaria.

Saltaia[69]

Gen. et sp. nov

Valid

Mergl & Herrera in Mergl et al.

Late Cambrian

Lampazar Formation

 Argentina

A member of Elkaniidae. The type species is Saltaia lampazarensis

Savagerhynchus[74]

Gen. et sp. nov

Valid

Baranov & Blodgett

Devonian (early Pragian)

 United States

A trigonirhynchiid rhynchonellid brachiopod. The type species is Savagerhynchus hecetaensis.

Scalenomena[82]

Gen. et comb. nov

Valid[83]

Cocks

Ordovician (early Katian

Craighead Limestone

 United Kingdom

A chilidiopsoidean brachiopod. A new genus for "Fardenia" scalena Williams (1962).

Schizambon cardonalis[69]

Sp. nov

Valid

Mergl & Herrera in Mergl et al.

Late Cambrian

Lampazar Formation

 Argentina

Schizobolus pilasiensis[85]

Sp. nov

Valid

Mergl in Morzadec et al.

Devonian (Givetian)

Floresta Formation

 Colombia

A member of the family Trematidae.

Sillakkudirhynchia[86]

Gen. et comb. nov

Valid

Radulović et al.

Late Cretaceous (Coniacian to early Maastrichtian)

Sillakkudi Formation

 India

A member of Rhynchonellida belonging to the family Tetrarhynchiidae. A new genus for "Rhynchonella" plicatiloides Stoliczka (1872); genus also contains "Cretirhynchia" sahnii Radulović & Ramamoorthy (1992).

Sivorthis precordillerana[61]

Sp. nov

Valid

Benedetto

Ordovician (Darriwilian)

Los Azules Formation
San Juan Formation

 Argentina

A member of Orthida belonging to the family Orthidae; a species of Sivorthis.

Talexirhynchia[87]

Gen. et sp. nov

Valid

Feldman et al.

Middle Jurassic (Callovian)

Mughanniyya Formation

 Jordan

A rhynchonellid brachiopod. The type species is Talexirhynchia kadishi.

Tethyochonetes minor[80]

Sp. nov

Valid

Zhang et al.

Permian (Changhsingian)

Longtan Formation

 China

A member of Productida belonging to the family Rugosochonetidae.

Thecocyrtella dagysii[56]

Sp. nov

Valid

Halamski et al.

Middle Triassic (Ladinian)

 Croatia

A member of Spiriferida; a species of Thecocyrtella.

Theodossia kagarmanovi[88]

Sp. nov

Valid

Bezgodova

Devonian (Frasnian)

 Russia

A member of Spiriferida belonging to the family Theodossiidae, a species of Theodossia.

Theodossia novosemelica[88]

Sp. nov

Valid

Bezgodova

Devonian (Frasnian)

 Russia

A member of Spiriferida belonging to the family Theodossiidae, a species of Theodossia.

Theodossia tcherkesovae[88]

Sp. nov

Valid

Bezgodova

Devonian (Frasnian)

 Russia

A member of Spiriferida belonging to the family Theodossiidae, a species of Theodossia.

Torobolus[77]

Gen. et sp. nov

Valid

Benedetto & Muñoz

Ordovician (Tremadocian)

Devendeus Formation

 Argentina

A member of Obolidae. The type species is Torobolus subplanus.

Trigonirhynchia celtiberica[64]

Sp. nov

Valid

GarcĂ­a-Alcalde & Herrera

Devonian (early Pragian)

 Spain

A member of Rhynchonellida belonging to the family Trigonirhynchiidae.

Trigonirhynchia palentina[64]

Sp. nov

Valid

GarcĂ­a-Alcalde & Herrera

Devonian (early Pragian)

 Spain

A member of Rhynchonellida belonging to the family Trigonirhynchiidae.

Yangtzeella chupananica[71]

Sp. nov

Valid

Popov, Holmer & Baars in Popov et al.

Ordovician (Darriwilian)

Chahgonbad Formation

 Iran

A clarkellid pentamerid brachiopod; a species of Yangtzeella.

Molluscs

Echinoderms

Name Novelty Status Authors Age Unit Location Notes Images

Aegyptiaris[89]

Gen. et sp. et comb. nov

Valid

Abdelhamid & Abdelghany

Cretaceous (Aptian to Cenomanian)

 Egypt

An emiratiid sea urchin. The type species is Aegyptiaris halalensis;[89] genus also contains "Trochodiadema" isidis Fourtau (1921).[90]

Aganaster jagiellonicus[91]

Sp. nov

Valid

Thuy, Kutscher & PƂachno

Carboniferous (late Tournaisian)

Mazurowe DoƂy Formation

 Poland

A brittle star belonging to the suborder Ophiurina, a species of Aganaster.

Archaeocidaris marmorcataractensis[92]

Sp. nov

Valid

Thompson et al.

Carboniferous (early Pennsylvanian)

Marble Falls Formation

 United States

A sea urchin, a species of Archaeocidaris.

Baficrinus[93]

Gen. et sp. nov

Valid

Prokop & NohejlovĂĄ

Devonian (late Emsian)

Daleje-Tƙebotov Formation

 Czech Republic

A crinoid belonging to the group Disparida and the family Zophocrinidae. The type species is Baficrinus vigilis.

Baudicrinus[94]

Gen. et sp. nov

Valid

Oji & Twitchett

Early Triassic

AI Jil Formation

 Oman

A dadocrinid articulate crinoid, a relative of encrinids. The type species is Baudicrinus krystyni.

Bizarroglobus[95]

Gen. et sp. nov

Valid

Sumrall & Sprinkle

Ordovician

Kanosh Shale

 United States

An edrioasteroid. The type species is Bizarroglobus medusae.

Browerocrinus[96]

Gen. et sp. nov

Valid

Ausich, Peter & Ettensohn

Silurian (Rhuddanian)

Brassfield Formation

 United States

A crinoid. The type species is Browerocrinus arthrikos.

Callocystites fresti[97]

Sp. nov

Valid

Paul

Silurian

Massie Formation

 United States

A callocystitid cystoid, a species of Callocystites.

‘Cidaris’ labergensis[98]

Sp. nov

Valid[99]

Zonneveld, Furlong & Sanders

Late Triassic

Aksala Formation

 Canada

A sea urchin. Zonneveld, Furlong & Sanders (2015) assign it to the form genus ‘Cidaris’, stressing that it does not imply affinity with the extant genus Cidaris.[98]

Conollia sporranoides[100]

Sp. nov

Valid

Rahman, Stewart & Zamora

Ordovician (Sandbian)

Balclatchie Formation

 United Kingdom

A member of Ctenocystoidea, a species of Conollia.

Deltoblastus beaveri[101]

Sp. nov

Valid

Morgan

Permian

Timor

A blastoid belonging to the family Schizoblastidae.

Deltoblastus elevatus[101]

Sp. nov

Valid

Morgan

Permian

Timor

A blastoid belonging to the family Schizoblastidae.

Deltoblastus ewini[101]

Sp. nov

Valid

Morgan

Permian

Timor

A blastoid belonging to the family Schizoblastidae.

Dimerocrinites (Dimerocrinites) cheilobathron[102]

Sp. nov

Valid

Clement & Brett

Early Devonian (Lochkovian)

Birdsong Shale

 United States

A camerate crinoid belonging to the group Diplobathrida.

Echinanthus callosensis[103]

Nom. nov

Valid

Carrasco

Eocene (Lutetian)

 Spain

A sea urchin; a replacement name for Echinanthus dorsalis Cotteau (1889).

Ekmelocrinus[104]

Gen. et comb. nov

Valid

Webster & Donovan

Permian

 Indonesia

A crinoid. Genus includes Ekmelocrinus amplior (Wanner, 1924), Ekmelocrinus subamplior (Wanner, 1949), Ekmelocrinus ovoides (Wanner, 1949), Ekmelocrinus verbeeki (Wanner, 1916) and Ekmelocrinus vermistriatus (Wanner, 1916).

Enakomusium[105]

Gen. et comb. nov

Valid

Thuy

Jurassic (Toarcian to Oxfordian)

 France
 Germany
  Switzerland
 United Kingdom

A brittle star. A new genus for "Ophioderma" weymouthiense Damon (1880); genus also contains "Ophiomusium" ferrugineum Boehm (1889), "Ophiolepis" leckenbyi Wright (1880), "Ophiomusium" geisingense Kutscher (1992), "Ophiomusium" mammillatum Hess (1966) and "Ophiura" gagnebini Thurmann (1851).

Eodevonocystis[102]

Gen. et sp. nov

Valid

Clement & Brett

Early Devonian (Lochkovian)

Birdsong Shale

 United States

A mitrate belonging to the group Anomalocystitida. The type species is E. marilynni.

Eodolatocrinus[102]

Gen. et sp. et comb. nov

Valid

Clement & Brett

Late Silurian-early Devonian

Bob Limestone (Brownsport Group)
Lower Birdsong Shale

 United States

A crinoid belonging to the group Monobathrida. The type species is E. hlabsei; genus also includes "Technocrinus" niagarensis Springer (1921).

Eohadroblastus[102]

Gen. et sp. nov

Valid

Clement & Brett

Early Devonian (Lochkovian)

Birdsong Shale

 United States

A blastoid belonging to the group Fissiculata. The type species is E. inexpectatus.

Eohalysiocrinus broweri[102]

Sp. nov

Valid

Clement & Brett

Early Devonian (Lochkovian)

Birdsong Shale
Rockhouse Limestone

 United States

A disparid crinoid belonging to the group Calceocrinida.

Eohalysiocrinus gibsoni[102]

Sp. nov

Valid

Clement & Brett

Early Devonian (Lochkovian)

Birdsong Shale
Rockhouse Limestone

 United States

A disparid crinoid belonging to the group Calceocrinida.

Eudimerocrinus hlabsei[102]

Sp. nov

Valid

Clement & Brett

Late Silurian (Pridoli)-early Devonian (Lochkovian)

Decatur Limestone
Rockhouse Limestone

 United States

A camerate crinoid belonging to the group Diplobathrida.

Exaesiodiscus derbyi[106]

Sp. nov

Valid

Scheffler, da Fonseca & Fernandes

Devonian (middle Eifelian)

Maecuru Formation

 Brazil

A crinoid, a species of Exaesiodiscus.

Frasericrinus[107]

Gen. et sp. nov

Valid

Ausich et al.

Early Cretaceous (probably early late Albian)

Peace River Formation

 Canada

An isocrinid crinoid. The type species is Frasericrinus mauricensis.

Globoeocrinus zhaoyuanlongensis[108]

Sp. nov

Valid

Yang et al.

Cambrian

Balang Formation

 China

An eocrinoid, a species of Globoeocrinus.

Guangxicrinus[109]

Gen. et sp. nov

Valid

Webster & Chen

Devonian (Eifelian)

Yingtang Formation

 China

A marsupiocrinid crinoid. The type species is Guangxicrinus xiangzhouensis.

Hemidiadema batalleri[110]

Sp. nov

Valid

Clément

Early Cretaceous (Aptian)

 Spain

A sea urchin belonging to the family Glyphocyphidae, a species of Hemidiadema.

Hemidiadema forcallensis[110]

Sp. nov

Valid

Clément

Early Cretaceous (Aptian)

 Spain

A sea urchin belonging to the family Glyphocyphidae, a species of Hemidiadema.

Hemidiadema malladai[110]

Sp. nov

Valid

Clément

Early Cretaceous (Aptian)

 Spain

A sea urchin belonging to the family Glyphocyphidae, a species of Hemidiadema.

Heviacrinus tromelini[111]

Sp. nov

Valid

Lefebvre et al.

Ordovician (late Darriwilian)

 France

A crinoid, a species of Heviacrinus.

Icthyocrinus erugatus[102]

Sp. nov

Valid

Clement & Brett

Early Devonian (Lochkovian)

Rockhouse Limestone

 United States

A cladid crinoid belonging to the group Sagenocrinida.

Iocrinus africanus[112]

Sp. nov

Valid

Zamora, Rahman & Ausich

Ordovician (Darriwilian)

Taddrist Formation

 Morocco

An iocrinid crinoid, a species of Iocrinus.

Lanxocolumnus (col.) chaleurensis[113]

Sp. nov

Valid

Donovan & Keighley

Silurian (Telychian)

 Canada

A crinoid, a species of Lanxocolumnus.

Lapidaster hellersi[114]

Sp. nov

Valid

Numberger-Thuy & Thuy

Middle Jurassic (earliest Bajocian)

 Luxembourg

A brittle star belonging to the family Ophiacanthidae, a species of Lapidaster.

Laudonomphalus ferreirai[106]

Sp. nov

Valid

Scheffler, da Fonseca & Fernandes

Devonian (middle Eifelian)

Maecuru Formation

 Brazil

A crinoid, a species of Laudonomphalus.

Llanocystis[115]

Gen. et sp. nov

Valid

Zamora, Sumrall & Sprinkle

Cambrian (Furongian)

Wilberns Formation

 United States

An eocrinoid of uncertain phylogenetic placement. The type species is Llanocystis wilbernsensis.

Lovenicystis kopfi[116]

Sp. nov

Valid

Paul

Silurian

 United States

A callocystitid cystoid, a species of Lovenicystis.

Marettocrinus hartti[106]

Sp. nov

Valid

Scheffler, da Fonseca & Fernandes

Devonian (middle Eifelian)

Maecuru Formation

 Brazil

A crinoid, a species of Marettocrinus.

Marettocrinus katzeri[106]

Sp. nov

Valid

Scheffler, da Fonseca & Fernandes

Devonian (middle Eifelian)

Maecuru Formation

 Brazil

A crinoid, a species of Marettocrinus.

Marginix[117]

Nom. nov

Valid

MartĂ­nez & del RĂ­o

Devonian

Talacasto Formation

 Argentina

A brittle star belonging to the order Oegophiurida and the family Encrinasteridae; a replacement name for Marginaster Haude (1995) and Marginura Haude (1999) (both preoccupied).

Macrostylocrinus tertibrachialis[102]

Sp. nov

Valid

Clement & Brett

Early Devonian (Lochkovian)

Rockhouse Limestone

 United States

A camerate crinoid belonging to the group Monobathrida.

Marsupiocrinus (Amarsupiocrinus) devonicus[102]

Sp. nov

Valid

Clement & Brett

Early Devonian (Lochkovian)

Birdsong Shale

 United States

A camerate crinoid belonging to the group Monobathrida.

Notatudiadema[89]

Gen. et sp. nov

Valid

Abdelhamid & Abdelghany

Late Cretaceous (Cenomanian)

 Egypt

A diadematoid sea urchin. The type species is Notatudiadema rekeibensis.

Ophiocantabria[118]

Gen. et sp. nov

Valid

Blake, Zamora & GarcĂ­a-Alcalde

Devonian (Lochkovian)

Furada Formation

 Spain

A brittle star belonging to the order Oegophiurida, suborder Lysophiurina and the family Encrinasteridae. The type species is Ophiocantabria elegans.

Ophiodoris reconciliator[105]

Sp. nov

Valid

Thuy

Middle Jurassic (early Bajocian)

Longwy Formation

 France

A brittle star belonging to the family Ophionereididae, a species of Ophiodoris.

Ophiotitanos aschmannicor[105]

Sp. nov

Valid

Thuy

Middle Jurassic (early Bajocian)

Longwy Formation

 France

A brittle star belonging to the family Ophiodermatidae, a species of Ophiotitanos.

Paracidaris (Anisocidaris) eluveitie[119]

Sp. nov

Valid

Hostettler & Menkveld-Gfeller

Late Jurassic (Oxfordian)

Wildegg Formation

  Switzerland

A member of Cidaroida, a species of Paracidaris.

Parahormocrinus[102]

Gen. et sp. nov

Valid

Clement & Brett

Late Silurian (Pridoli)

Decatur Limestone

 United States

A cladid crinoid belonging to the group Sagenocrinida. The type species is P. decaturensis.

Paramarsupiocrinus[102]

Gen. et sp. nov

Valid

Clement & Brett

Late Silurian (Pridoli)

 United States

A camerate crinoid belonging to the group Monobathrida. The type species is P. broadheadi.

Parapatelliocrinus[102]

Gen. et sp. nov

Valid

Clement & Brett

Devonian (Lochkovian)

Lower Rockhouse Limestone

 United States

A crinoid belonging to the group Monobathrida. The type species is P. broweri.

Pentaridica mendesi[106]

Sp. nov

Valid

Scheffler, da Fonseca & Fernandes

Devonian (middle Eifelian)

Maecuru Formation

 Brazil

A crinoid, a species of Pentaridica.

Polycidaris gygii[119]

Sp. nov

Valid

Hostettler & Menkveld-Gfeller

Late Jurassic (Oxfordian)

Lochen Formation
Wildegg Formation

 Germany
  Switzerland

A member of Cidaroida, a species of Polycidaris.

Porosublastus[120]

Gen. et sp. nov

Valid

Sprinkle & Sumrall

Early Ordovician

Ninemile Shale

 United States

An edrioasteroid. The type species is Porosublastus inexpectus.

Probalocrinus[102]

Gen. et comb. nov

Valid

Clement & Brett

Late Silurian (Ludlovian)-early Devonian (Lochkovian)

Birdsong Shale
Henryhouse Formation

 United States

A camerate crinoid belonging to the group Diplobathrida. The type species is "Siphonocrinus" dignis Strimple (1963).

Protogloboeocrinus[121]

Gen. et sp. nov

Valid

Zhao et al.

Cambrian

Balang Formation

 China

A member of Eocrinoidea. Genus includes new species P. yini Zhao, Peng & Wu in Zhao et al. (2015).

Pseudedriophus[120]

Gen. et sp. nov

Valid

Sprinkle & Sumrall

Early Ordovician

Ninemile Shale

 United States

An edrioasteroid. The type species is Pseudedriophus guensburgi.

Quechuacrinus[122]

Gen. et sp. nov

Valid

Guensburg & Waisfeld

Ordovician (Floian)

 Argentina

A crinoid. The type species is Quechuacrinus ticsa.

Ramseyocrinus argentinus[122]

Sp. nov

Valid

Guensburg & Waisfeld

Ordovician (Floian)

 Argentina

A crinoid, a species of Ramseyocrinus.

Rautscholdticrinus[104]

Gen. et comb. nov

Valid

Webster & Donovan

Permian

 Indonesia

A crinoid. Genus includes Rautscholdticrinus indicus (Wanner, 1916) and Rautscholdticrinus weidnerii (Wanner, 1937).

Rhabdocidaris deubelbeissi[119]

Sp. nov

Valid

Hostettler & Menkveld-Gfeller

Late Jurassic (Oxfordian)

Wildegg Formation

  Switzerland

A member of Cidaroida, a species of Rhabdocidaris.

Ristnacrinus oepiki[123]

Sp. nov

Valid

Ausich, Rozhnov & Kammer

Ordovician (Katian)

 Estonia

A crinoid related to Iocrinus, a species of Ristnacrinus.

Salamonicrinus[124]

Gen. et sp. nov

Valid

Trzęsiok

Late Jurassic (Tithonian)

 Poland

A cyrtocrinid crinoid. The type species is Salamonicrinus prodigiosum.

Salirocystis[116]

Gen. et comb. nov

Valid

Paul

Silurian

 United States

A callocystitid cystoid; a new genus for "Apiocystites" anna Safford (1869).

Temnocrinus americanus[96]

Sp. nov

Valid

Ausich, Peter & Ettensohn

Silurian (Rhuddanian)

Brassfield Formation

 United States

A crinoid, a species of Temnocrinus.

Velocrinus[125]

Gen. et sp. nov

Valid

Ausich, Wilson & Vinn

Late Silurian

Kaugatuma Formation

 Estonia

A crotalocrinitid crinoid. The type species is Velocrinus coniculus.

Conodonts

Name Novelty Status Authors Age Unit Location Notes Images

Arianagnathus[126]

Gen. et sp. nov

Valid

MĂ€nnik, Miller & Hairapetian

Silurian

Niur Formation

 Iran

A member of Prioniodontida, possibly a balognathid. The type species is Arianagnathus jafariani.

Dichognathus zhaoi[127]

Sp. nov

Valid

Xiong, Wu & Wang

Ordovician

Tarim Basin

 China

Eolinguipolygnathus radula[128]

Sp. nov

Valid

Aboussalam & Becker in Aboussalam, Becker & Bultynck

Devonian (early Emsian)

 Morocco

Eoplacognathus aisangensis[127]

Sp. nov

Valid

Xiong, Wu & Wang

Ordovician

Tarim Basin

 China

Gapparodus gapparites[129]

Sp. nov

Valid

Yang, Steiner & Keupp

Early Cambrian

Shuijingtuo Formation

 China

A member of Furnishinidae; a species of Gapparodus.

Hayashiella[130]

Gen. et comb. nov

Valid

Kiliç et al.

Late Triassic

 Canada
 Italy
 Japan
 Slovakia

A member of Gondolellidae. A new genus for "Metapolygnathus" nodosus Hayashi (1968); genus also includes "Gondolella" carpathica Mock (1979), H. lindae (Orchard) and H. permica (Hayashi).

Hindeodus capitanensis[131]

Sp. nov

Valid

Wardlaw & Nestell

Permian

 United States

Icriodus ovalis[128]

Sp. nov

Valid

Bultynck in Aboussalam, Becker & Bultynck

Devonian (late Emsian)

MdĂąouer-el-KbĂźr Formation

 Morocco

Icriodus praerectirostratus[128]

Sp. nov

Valid

Bultynck in Aboussalam, Becker & Bultynck

Devonian (late Emsian)

Timrhanrhart Formation

 Morocco

Mazzaella[130]

Gen. et comb. nov

Valid

Kiliç et al.

Late Triassic

 Germany
 Hungary
 Italy
 Turkey

A member of Gondolellidae. A new genus for "Epigondolella" carnica Krystyn (1975); genus also includes "Metapolygnathus" baloghi Kovacs (1977).

Moskalenkodus[132]

Gen. et comb. nov

Valid

Dzik

Late Ordovician

Krivaya Luka Formation
Makarovo Formation
Tunguz Basin

 Russia

A member of Prioniodontida belonging to the family Distomodontidae. A new genus for "Amorphognathus" quinquiradiatus Moskalenko in Kanygin et al. (1977). Genus also includes Moskalenkodus cruciformis (Moskalenko, 1970).

Neostreptognathodus costatus[133]

Sp. nov

Valid

Wang & Zhang

Early Permian

Amushan Formation

 China

Ozarkodina derenjalensis[126]

Sp. nov

Valid

MĂ€nnik, Miller & Hairapetian

Silurian

Niur Formation

 Iran

A member of Ozarkodinida, a spathognathodontid; a species of Ozarkodina.

Polygnathus bicristatus[134]

Sp. nov

Valid

Mossoni et al.

Late Devonian or Early Carboniferous

 Italy

A member of Ozarkodinida belonging to the family Polygnathidae, a species of Polygnathus.

Polygnathus nuragicus[134]

Sp. nov

Valid

Mossoni et al.

Late Devonian or Early Carboniferous

 Italy

A member of Ozarkodinida belonging to the family Polygnathidae, a species of Polygnathus.

Postoistodus[127]

Gen. et 2 sp. nov

Valid

Xiong, Wu & Wang

Ordovician

Tarim Basin

 China

Genus includes new species P. elongatus and P. tridens.

Pseudohindeodus brevis[131]

Sp. nov

Valid

Wardlaw & Nestell

Permian

 United States

Pseudopolygnathus granulobatus[134]

Sp. nov

Valid

Mossoni et al.

Late Devonian or Early Carboniferous

 Italy

A member of Ozarkodinida belonging to the family Polygnathidae, a species of Pseudopolygnathus.

Pygodus serrus ambonis[127]

Subsp. nov

Valid

Xiong, Wu & Wang

Ordovician

Tarim Basin

 China

Spathogondolella[135]

Gen. et sp. nov

Valid

Jiang in Chen et al.

Early Triassic

 China

The type species is Spathogondolella jiarongensis.

Vjalovognathus nicolli[136]

Sp. nov

Valid[137]

Yuan, Shen & Henderson in Yuan et al.

Early Permian

Xiala Formation

 China

A species of Vjalovognathus.

Fishes

Amphibians

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Eocyclotosaurus appetolatus[141]

Sp. nov

Valid

Rinehart, Lucas & Schoch

Middle Triassic

Moenkopi Formation

 United States

A capitosaur, a species of Eocyclotosaurus.

Metoposaurus algarvensis[142]

Sp. nov

Valid

Brusatte et al.

Late Triassic

 Portugal

A species of Metoposaurus.

Parapytanga[143]

Gen. et sp. nov

Valid

Strapasson, Pinheiro & Soares

Middle or late Permian

Rio do Rasto Formation

 Brazil

A basal member of Stereospondylomorpha. The type species is Parapytanga catarinensis.

Procuhy[144]

Gen. et sp. nov

Valid

Cisneros et al.

Permian (Cisuralian)

Lower Pedra de Fogo Formation

 Brazil

A member of Trimerorhachidae. The type species is Procuhy nazariensis.

Timonya[144]

Gen. et sp. nov

Valid

Cisneros et al.

Permian (Cisuralian)

Lower Pedra de Fogo Formation

 Brazil

A member of Dvinosauria. The type species is Timonya anneae.

Name Novelty Status Authors Age Unit Location Notes Images

Aletrimyti[145]

Gen. et sp. nov

Valid

Szostakiwskyj, Pardo & Anderson

Permian (Kungurian)

Fairmont Shale

 United States

A member of Recumbirostra. The type species is Aletrimyti gaskillae.

Batropetes appelensis[146]

Sp. nov

Valid

Glienke

Early Permian

Saar–Nahe Basin

 Germany

A species of Batropetes.

Batropetes palatinus[146]

Sp. nov

Valid

Glienke

Early Permian

Saar–Nahe Basin

 Germany

A species of Batropetes.

Dvellecanus[145]

Gen. et sp. nov

Valid

Szostakiwskyj, Pardo & Anderson

Permian (Kungurian)

Fairmont Shale

 United States

A member of Recumbirostra. The type species is Dvellecanus carrolli.

Name Novelty Status Authors Age Unit Location Notes Images

Bufo mirus[147]

Sp. nov

Valid

Syromyatnikova

Late Early – early Middle Miocene

Khalagay Formation

 Russia

A true toad of uncertain phylogenetic placement, most similar and possibly related to the Mongolian toad.

Hensonbatrachus[148]

Gen. et sp. nov

Valid

Gardner & Brinkman

Late Cretaceous (Campanian)

Dinosaur Park Formation
Oldman Formation

 Canada

A frog. The type species is Hensonbatrachus kermiti.

Lepidobatrachus australis[149]

Sp. nov

Valid

Nicoli

Late Miocene to Early Pliocene

Monte Hermoso Formation

 Argentina

A frog belonging to the family Ceratophryidae, a species of Lepidobatrachus.

Palaeoplethodon[150]

Gen. et sp. nov

Valid

Poinar & Wake

Eocene or Miocene

 Dominican Republic

A lungless salamander found in Dominican amber. The type species is Palaeoplethodon hispaniolae.

Tyrrellbatrachus[151]

Gen. et sp. nov

Valid

Gardner

Late Cretaceous (Campanian)

Dinosaur Park Formation

 Canada

A frog of uncertain phylogenetic placement. The type species is Tyrrellbatrachus brinkmani.

Varibatrachus[152]

Gen. et sp. nov

Valid

Parmley, Chandler & Chandler

Miocene (late Clarendonian)

Ash Hollow Formation

 United States

A frog of uncertain phylogenetic placement. The type species is Varibatrachus abraczinskasae.

Name Novelty Status Authors Age Unit Location Notes Images

Alveusdectes[153]

Gen. et sp. nov

Valid

Liu & Bever

Late Permian (Wuchiapingian, roughly 256 Ma)

Shangshihezi Formation

 China

A diadectid. The type species is Alveusdectes fenestralis.

Ichthyosauromorphs

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Cartorhynchus[156]

Gen. et sp. nov

Valid

Motani et al.

Early Triassic (Olenekian)

Nanlinghu Formation

 China

A relative of ichthyopterygians. The type species is Cartorhynchus lenticarpus.

Dearcmhara[157]

Gen. et sp. nov

Valid

Brusatte et al.

Jurassic (Toarcian–Bajocian)

 United Kingdom

A basal member of Neoichthyosauria. The type species is Dearcmhara shawcrossi.

Eretmorhipis[158]

Gen. et sp. nov

Valid

Chen et al.

Early Triassic (late Spathian)

Jialingjiang Formation

 China

A hupehsuchian. The type species is Eretmorhipis carrolldongi.

Ichthyosaurus anningae[159]

Sp. nov

Valid

Lomax & Massare

Early Jurassic (Hettangian/Sinemurian–Pliensbachian)

 United Kingdom

A species of Ichthyosaurus.

Muiscasaurus[160]

Gen. et sp. nov

Valid[161]

Maxwell et al.

Early Cretaceous (Barremian–Aptian)

Paja Formation

 Colombia

A member of Ophthalmosauridae. The type species is Muiscasaurus catheti.

Sauropterygians

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Anguanax[165]

Gen. et sp. nov

Valid[166]

Cau & Fanti

Late Jurassic (Oxfordian)

Rosso Ammonitico Veronese Formation

 Italy

A pliosaurid. The type species is Anguanax zignoi.

Atychodracon[167]

Gen. et comb. nov

Valid

Smith

Early Jurassic

 United Kingdom

A rhomaleosaurid plesiosaur; a new genus for "Plesiosaurus" megacephalus Stutchbury (1846).

Cardiocorax[168]

Gen. et sp. nov

Valid

AraĂșjo et al.

Late Cretaceous (early Maastrichtian)

Mocuio Formation

 Angola

An elasmosaurid plesiosaur. The type species is Cardiocorax mukulu.

Dianmeisaurus[169]

Gen. et sp. nov

Valid

Shang & Li

Middle Triassic (Anisian)

Guangling Formation

 China

A member of Eosauropterygia of uncertain phylogenetic placement, more closely related to nothosaurs than to plesiosaurs. The type species is Dianmeisaurus gracilis.

Makhaira[170]

Gen. et sp. nov

Valid

Fischer et al.

Early Cretaceous (Hauterivian)

 Russia

A pliosaurid. The type species is Makhaira rossica.

Vegasaurus[171]

Gen. et sp. nov

Valid

O’Gorman et al.

Late Cretaceous (early Maastrichtian)

Snow Hill Island Formation

Antarctica (Vega Island)

An elasmosaurid plesiosaur. The type species is Vegasaurus molyi.

Wangosaurus[172]

Gen. et sp. nov

Valid

Ma et al.

Middle Triassic (Ladinian)

Falang Formation

 China

A pistosauroid. The type species is Wangosaurus brevirostris.

Lepidosaurs

Research

  • A phylogenetic analysis of living and fossil squamate relationships, based on morphological and molecular data, is published by Reeder et al. (2015).[173]
  • Miocene anoles from the Dominican amber, showing the habitat specializations also present in the extant Caribbean anoles, are described by Sherratt et al. (2015).[174]

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Clevosaurus sectumsemper[175]

Sp. nov

Valid

Klein et al.

Late Triassic (Rhaetian)

 United Kingdom

A clevosaurid sphenodontian, a species of Clevosaurus.

Name Novelty Status Authors Age Unit Location Notes Images

Archaerhineura[176]

Gen. et sp. nov

Valid

Longrich et al.

Late Palaeocene

Polecat Bench Formation

 United States

A rhineurid amphisbaenian. The type species is Archaerhineura mephitis.

Asagaolacerta[177]

Gen. et sp. nov

Valid

Evans & Matsumoto

Early Cretaceous

Kuwajima Formation

 Japan

A lizard of uncertain phylogenetic placement. The type species is Asagaolacerta tricuspidens.

Asprosaurus[178]

Gen. et sp. nov

Valid

Park, Evans & Huh

Late Cretaceous

 South Korea

A member of Anguimorpha, probably a member of Monstersauria. The type species is Asprosaurus bibongriensis.

Babibasiliscus[179]

Gen. et sp. nov

Valid

Conrad

Eocene (approximately 48 Ma)

Bridger Formation

 United States

A member of Corytophanidae. The type species is Babibasiliscus alxi.

Barbatteius[180]

Gen. et sp. nov.

Valid[181]

Venczel & Codrea

Late Cretaceous (early Maastrichtian)

HaĆŁeg Basin

 Romania

Originally assigned to the family Teiidae, but subsequently transferred to the separate family Barbatteiidae within the group Teiioidea.[182] The type species is Barbatteius vremiri.

Cadurcogekko verus[183]

Sp. nov

Valid

Bolet et al.

Eocene

 France

A gekkotan lizard, a species of Cadurcogekko.

Chromatogenys[184]

Gen. et sp. nov

Valid

MakĂĄdi & Nydam

Late Cretaceous (Santonian)

 Hungary

A member of Scincomorpha of uncertain phylogenetic placement. The type species is Chromatogenys tiliquoides.

Chthonophis[176]

Gen. et sp. nov

Valid

Longrich et al.

Probably early Palaeocene

Bug Creek Anthills, Fort Union Formation

 United States

An amphisbaenian, the only member of the new family Chthonophidae. The type species is Chthonophis subterraneus.

Cuvieribaena[185]

Gen. et sp. nov

Valid

ČerƈanskĂœ, AugĂ©c & Rage

Eocene (Bartonian)

 France

A blanid amphisbaenian. The type species is Cuvieribaena carlgansi.

Dryadissector[186]

Gen. et sp. nov

Valid

Wick, Lehman & Brink

Late Cretaceous (early Campanian)

Aguja Formation

 United States

A member of Varanoidea. The type species is Dryadissector shilleri.

Gekkomimus[183]

Gen. et comb. nov

Valid

Bolet et al.

Eocene

 France

A skink; a new genus for "Cadurcogekko" rugosus Augé (2005).

Gueragama[187]

Gen. et sp. nov

Valid

SimÔes et al.

Late Cretaceous (Turonian-Campanian)

Goio-ErĂȘ Formation

 Brazil

The first member of Acrodonta (a relative of agamids and chameleons) described from South America. The type species is Gueragama sulamericana.

Hakuseps[177]

Gen. et sp. nov

Valid

Evans & Matsumoto

Early Cretaceous

 Japan

A squamate of uncertain phylogenetic placement. The type species is Hakuseps imberis.

Kuroyuriella[177]

Gen. et sp. nov

Valid

Evans & Matsumoto

Early Cretaceous

Kuwajima Formation

 Japan

A lizard of uncertain phylogenetic placement. The type species is Kuroyuriella mikikoi.

Norellius[188]

Gen. et sp. nov

Valid

Conrad & Daza

Early Cretaceous (probably approximately 130 Mya)

Öösh Basin

 Mongolia

A member of Gekkonomorpha of uncertain phylogenetic placement. The type species is Norellius nyctisaurops.

Ophisaurus holeci[189]

Sp. nov

Valid

Klembara

Miocene

 Czech Republic
 Germany[190][191]

A glass lizard.

Pachygenys adachii[192]

Sp. nov

Valid

Ikeda, Ota & Saegusa

Early Cretaceous

Sasayama Group

 Japan

A squamate of uncertain phylogenetic placement, a species of Pachygenys.

Phosphorosaurus ponpetelegans[193]

Sp. nov

Valid[194]

Konishi et al.

Late Cretaceous (Maastrichtian)

 Japan

A halisaurine mosasaur, a species of Phosphorosaurus.

Plioplatecarpus peckensis[195]

Sp. nov

Valid

Cuthbertson & Holmes

Late Cretaceous (Campanian)

Bearpaw Formation

 United States

A mosasaur, a species of Plioplatecarpus.

Pseudopus rugosus[189]

Sp. nov

Valid

Klembara

Early Miocene

 Czech Republic

A relative of the sheltopusik.

Tetrapodophis[196]

Gen. et sp. nov

Valid

Martill, Tischlinger & Longrich

Early Cretaceous (Aptian)

Crato Formation

 Brazil

A squamate reptile of uncertain phylogenetic placement; originally classified as an early, four-legged snake,[196] but subsequently argued to be a dolichosaurid.[197][198] The type species is Tetrapodophis amplectus.

Name Novelty Status Authors Age Unit Location Notes Images

Diablophis[199]

Gen. et comb. nov

Valid

Caldwell et al.

Late Jurassic

Morrison Formation

 United States

A basal snake; a new genus for "Parviraptor" gilmorei Evans (1996).

Eophis[199]

Gen. et sp. nov

Valid

Caldwell et al.

Middle Jurassic (Bathonian)

Forest Marble

 United Kingdom

A basal snake. The type species is Eophis underwoodi.

Portugalophis[199]

Gen. et sp. nov

Valid

Caldwell et al.

Late Jurassic (Kimmeridgian)

Camadas de Guimarota

 Portugal

A basal snake. The type species is Portugalophis lignites.

Renenutet[200]

Gen. et sp. nov

Valid

Mccartney & Seiffert

Eocene (Priabonian)

 Egypt

A member of Colubroidea. The type species is Renenutet enmerwer.

Turtles

Name Novelty Status Authors Age Unit Location Notes Images

Adocus sengokuensis[201]

Sp. nov

Valid

Sonoda et al.

Early Cretaceous

Sengoku Formation

 Japan

A species of Adocus.

Arvinachelys[202]

Gen. et sp. nov

Valid

Lively

Late Cretaceous (Campanian)

Kaiparowits Formation

 United States

A member of Baenidae. The type species is Arvinachelys goldeni.

Bairdemys thalassica[203]

Sp. nov

Valid

Ferreira et al.

Miocene

Capadare Formation

 Venezuela

A podocnemidid belonging to the clade Stereogenyina, a species of Bairdemys.

Baltemys velogastros[204]

Sp. nov

Valid

Lichtig & Lucas

Eocene (early Bridgerian)

Huerfano Formation

 United States

A member of Kinosternidae, a species of Baltemys.

Corsochelys bentleyi[205]

Sp. nov

Valid

Schwimmer et al.

Late Cretaceous

 United States

A member of Dermochelyidae, a species of Corsochelys.

Desmatochelys padillai[206]

Sp. nov

Valid

Cadena & Parham

Early Cretaceous (late Barremian-early Aptian)

Paja Formation

 Colombia

A member of Protostegidae, species of Desmatochelys.

Gaffneylania[207]

Gen. et sp. nov

Valid

Sterli, de la Fuente & Krause

Middle Eocene

Sarmiento Formation

 Argentina

A member of Meiolaniidae. The type species is Gaffneylania auricularis.

Gobiapalone palaeocenica[208]

Sp. nov

Valid

Danilov et al.

Paleocene

Bugin Tsav Basin

 Mongolia

A member of Trionychidae, a species of Gobiapalone. The genus Gobiapalone was considered to be a junior synonym of the genus Kuhnemys by Georgalis & Joyce (2017), though the authors maintained G. palaeocenica as a distinct species within the latter genus.[209]

Gomphochelys[210]

Gen. et sp. nov

Valid

Bourque et al.

Early Wasatchian

Willwood Formation

 United States

A member of Dermatemydidae. The type species is Gomphochelys nanus.

Judithemys russelli[211]

Sp. nov

Valid

Brinkman

Late Cretaceous

 Canada

A member of (likely non-monophyletic) Macrobaenidae, a species of Judithemys.

Jurassichelon[212]

Gen. et sp. nov

Valid

PĂ©rez-GarcĂ­a

Late Jurassic (early Tithonian)

 France

A basal member of Eucryptodira. The type species is Jurassichelon oleronensis.

Khunnuchelys lophorhothon[213]

Sp. nov

Valid

Danilov et al.

Late Cretaceous (Santonian or early Campanian)

Bostobe Formation

 Kazakhstan

A trionychine trionychid, a species of Khunnuchelys.

Kimurachelys[214]

Gen. et sp. nov

Valid

Brinkman et al.

Late Cretaceous (late Campanian)

Dinosaur Park Formation

 Canada

A chelonioid turtle. The type species is Kimurachelys slobodae.

Leyvachelys[215]

Gen. et sp. nov

Valid

Cadena

Early Cretaceous (Barremian-Aptian)

Glen Rose Formation
Paja Formation

 Colombia
 United States

A member of Pan-Cryptodira belonging to the family Sandownidae. The type species is Leyvachelys cipadi.

Mauremys oshiroi[216]

Sp. nov.

Valid

Takahashi et al.

Late Pleistocene

Tomori Amaga Cave

 Japan

A pond turtle.

Neurankylus hutchisoni[217]

Sp. nov

Valid[218]

Lively

Late Cretaceous (Campanian)

Kaiparowits Formation

 United States

A member of Baenidae, a species of Neurankylus.

Neurankylus utahensis[217]

Sp. nov

Valid[218]

Lively

Late Cretaceous (Campanian)

Kaiparowits Formation

 United States

A member of Baenidae, a species of Neurankylus.

Perochelys[219]

Gen. et sp. nov.

Valid

Li, Joyce & Liu

Early Cretaceous (Aptian)

Jiufotang Formation

 China

A soft-shelled turtle of uncertain phylogenetic placement; it might be a stem- or a crown-trionychid. The type species is Perochelys lamadongensis.

Phunoichelys[220]

Gen. et sp. nov.

Valid

Tong et al.

Late Jurassic

Phu Kradung Formation

 Thailand

A xinjiangchelyid eucryptodiran. The type species is Phunoichelys thirakhupti.

Portlandemys gracilis[221]

Sp. nov.

Valid

Anquetin, PĂŒntener & Billon-Bruyat

Late Jurassic (late Kimmeridgian)

Reuchenette Formation

  Switzerland

A plesiochelyid eucryptodiran, a species of Portlandemys.

Sternotherus bonevalleyensis[222]

Sp. nov.

Valid

Bourque & Schubert

Late Hemphillian

 United States

A musk turtle.

Sternotherus palaeodorus[222]

Sp. nov.

Valid

Bourque & Schubert

Late Hemphillian

 United States

A musk turtle.

Taraschelon[223]

Gen. et comb. nov

Valid[224]

PĂ©rez-GarcĂ­a

Early Oligocene

 France

A tortoise; a new genus for "Testudo" gigas Bravard (1844).

Testudo brevitesta[225]

Sp. nov

Valid[226]

Vlachos & Tsoukala

Late Pliocene (early Villafranchian)

 Greece

A tortoise, a species of Testudo.

Thalassemys bruntrutana[227]

Sp. nov

Valid

PĂŒntener, Anquetin & Billon-Bruyat

Late Jurassic (late Kimmeridgian)

Reuchenette Formation

  Switzerland

A basal member of Pancryptodira, a species of Thalassemys.

Toremys[228]

Gen. et sp. nov.

Valid

PĂ©rez-GarcĂ­a et al.

Early Cretaceous (early Albian)

 Spain

A pleurosternid paracryptodiran. The type species is Toremys cassiopeia.

"Trionyx" jixiensis[229]

Sp. nov.

Valid

Li et al.

Early Cretaceous

Chengzihe Formation

 China

A member of Trionychinae of uncertain phylogenetic placement.

Xiaochelys[230]

Gen. et sp. nov

Valid

Zhou & Rabi

Early Cretaceous (Barremian-early Aptian)

Yixian Formation

 China

A member of Sinemydidae. The type species is Xiaochelys ningchengensis.

Archosauriformes

Pseudosuchians

  • Revision of the type material of Rauisuchus tiradentes is published by Lautenschlager and Rauhut (2015).[231]
  • A study on the changes in the morphology of vertebrae caused by increased adaptation to aquatic locomotion in the evolution of crocodylomorphs is published by Molnar et al. (2015).[232]
  • A study of the evolutionary history of the clade Crocodyliformes is published by Bronzati, Montefeltro and Langer (2015).[233]
  • A study of impact of the climate changes on the evolution and biodiversity of pseudosuchian archosaurs is published by Mannion et al. (2015).[234]
  • A study on the phylogenetic position of thalattosuchians within Crocodylomorpha is published by Wilberg (2015).[235]
  • A review of the diagnostic features of the species assigned to the genus Machimosaurus is published by Martin, Vincent & Falconnet (2015).[236]
  • An isolated metriorhynchid tooth is described from the Early Cretaceous (Aptian) Hybla Formation (Sicily, Italy) by Chiarenza et al. (2015), extending the known geological range of Metriorhynchidae and Thalattosuchia by approximately 7–8 million years;[237] Fischer et al. (2015) subsequently caution that it might potentially be a pliosaurid tooth instead.[170]
Name Novelty Status Authors Age Unit Location Notes Images

Albertosuchus[238]

Gen. et sp. nov

Valid

Wu & Brinkman

Latest Cretaceous

Scollard Formation

 Canada

A crocodylian, probably a crocodyloid. The type species is Albertosuchus knudsenii.

Allodaposuchus hulki[239]

Sp. nov

Valid

Blanco et al.

Late Cretaceous (Maastrichtian)

Tremp Formation

 Spain

A eusuchian crocodylomorph, a species of Allodaposuchus.

Caiman wannlangstoni[240]

Sp. nov

Valid

Salas-Gismondi et al.

Late Middle Miocene

Pebas Formation

 Peru

A caiman, a species of Caiman.

Carnufex[241]

Gen. et sp. nov

Valid

Zanno et al.

Late Triassic (Carnian)

Pekin Formation

 United States

An early member of Crocodylomorpha. The type species is Carnufex carolinensis.

Dagasuchus[242]

Gen. et sp. nov

Valid

Lacerda, Schultz & Bertoni-Machado

Triassic (late Ladinian/early Carnian)

Santa Maria Formation

 Brazil

A non-crocodylomorph loricatan. The type species is Dagasuchus santacruzensis.

Gnatusuchus[240]

Gen. et sp. nov

Valid

Salas-Gismondi et al.

Late Middle Miocene

Pebas Formation

 Peru

A caiman. The type species is Gnatusuchus pebasensis.

Gorgetosuchus[243]

Gen. et sp. nov

Valid

Heckert et al.

Late Triassic

Pekin Formation

 United States

An aetosaur. The type species is Gorgetosuchus pekinensis.

Koumpiodontosuchus[244]

Gen. et sp. nov

Valid

Sweetman, Pedreira-Segade & Vidovic

Early Cretaceous (Barremian)

Wessex Formation

 United Kingdom

A crocodyliform related to Bernissartia. The type species is Koumpiodontosuchus aprosdokiti.

Kuttanacaiman[240]

Gen. et sp. nov

Valid

Salas-Gismondi et al.

Late Middle Miocene

Pebas Formation

 Peru

A caiman. The type species is Kuttanacaiman iquitosensis.

Lohuecosuchus[245]

Gen. et 2 sp. nov

Valid

NarvĂĄez et al.

Late Cretaceous (late Campanian-early Maastrichtian)

 France
 Spain

A eusuchian crocodyliform related to Allodaposuchus. The type species is Lohuecosuchus megadontos; genus also contains Lohuecosuchus mechinorum.

Zoneait[246]

Gen. et sp. nov

Valid

Wilberg

Middle Jurassic (Aalenian–Bajocian)

Snowshoe Formation

 United States

A thalattosuchian crocodylomorph. The type species is Zoneait nargorum.

Non-avian dinosaurs

Name Novelty Status Authors Age Unit Location Notes Images

Augustynolophus[253]

Gen. et comb. nov

Valid

Prieto-MĂĄrquez et al.

Late Cretaceous (late Maastrichtian)

Moreno Formation

 United States

A hadrosaurid; a new genus for "Saurolophus" morrisi Prieto-MĂĄrquez & Wagner (2013).

Boreonykus[254]

Gen. et sp. nov

Valid

Bell & Currie

Late Cretaceous (late Campanian)

Wapiti Formation

 Canada

A dromaeosaurid theropod. The type species is Boreonykus certekorum.

Chilesaurus[255]

Gen. et sp. nov.

Valid

Novas et al.

Late Jurassic (Tithonian)

Toqui Formation

 Chile

A dinosaur of uncertain phylogenetic placement. Originally classified as a theropod dinosaur (a basal member of Tetanurae), but subsequently argued to be an ornithischian.[256] The type species is Chilesaurus diegosuarezi.

Crichtonpelta[257]

Gen. et comb. nov

Valid[258]

Arbour & Currie

Late Cretaceous (Cenomanian–Turonian)

Sunjiawan Formation

 China

An ankylosaurine ankylosaurid; a new genus for "Crichtonsaurus" benxiensis LĂŒ, Ji, Gao & Li (2007).

Dakotaraptor[259]

Gen. et sp. nov.

Valid

DePalma et al.

Late Cretaceous (Maastrichtian)

Hell Creek Formation

 United States

A member of Dromaeosauridae. The type species is Dakotaraptor steini.

Eucnemesaurus entaxonis[260]

Sp. nov

Valid

McPhee et al.

Late Triassic

Lower Elliot Formation

 South Africa

A non-sauropod sauropodomorph, a species of Eucnemesaurus.

Galeamopus[247]

Gen. et comb. nov

Valid

Tschopp, Mateus & Benson

Late Jurassic (Kimmeridgian)

Morrison Formation

 United States

A diplodocid sauropod; a new genus for "Diplodocus" hayi Holland (1924).

Haestasaurus [261]

Gen. et comb. nov.

Valid

Upchurch, Mannion & Taylor

Early Cretaceous (late Berriasian—Valanginian)

Hastings Beds

 United Kingdom

A macronarian sauropod of uncertain phylogenetic placement; a new genus for "Pelorosaurus" becklesii Mantell (1852).

Horshamosaurus[262]

Gen. et comb. nov

Valid

Blows

Early Cretaceous

 United Kingdom

A new genus for "Polacanthus" rudgwickensis Blows (1996).

Hualianceratops [263]

Gen. et sp. nov.

Valid

Han et al.

Late Jurassic (Oxfordian)

Shishugou Formation

 China

A basal ceratopsian, possibly a member of Chaoyangsauridae. The type species is Hualianceratops wucaiwanensis.

Huanansaurus[264]

Gen. et sp. nov.

Valid

LĂŒ et al.

Late Cretaceous (Campanian-Maastrichtian)

Nanxiong Formation

 China

An oviraptorid theropod. The type species is Huanansaurus ganzhouensis.

Iguanodon galvensis[265]

Sp. nov

Valid

VerdĂș et al.

Early Cretaceous (early Barremian)

Galve, Teruel

 Spain

An ornithopod, a species of Iguanodon.

Ischioceratops[266]

Gen. et sp. nov.

Valid

He et al.

Late Cretaceous

Wangshi Group

 China

A leptoceratopsid ceratopsian. The type species is Ischioceratops zhuchengensis.

Koshisaurus[267]

Gen. et sp. nov

Valid

Shibata & Azuma

Early Cretaceous

Kitadani Formation

 Japan

A non-hadrosaurid hadrosauroid. The type species is Koshisaurus katsuyama.

Kunbarrasaurus[268]

Gen. et sp. nov.

Valid

Leahey et al.

Cretaceous (late Albian–(?)early Cenomanian)

Allaru Mudstone

 Australia

An ankylosaur. The type species is Kunbarrasaurus ieversi.

Lepidocheirosaurus[269]

Gen. et sp. nov.

Valid

Alifanov & Saveliev

Late Jurassic (Tithonian?)

Ukureyskaya Formation

 Russia

A dinosaur of uncertain phylogenetic placement; considered to be a theropod dinosaur related to Nqwebasaurus by the authors of its description,[269] but subsequently argued to be an ornithischian, possibly synonymous with Kulindadromeus.[270] The type species is Lepidocheirosaurus natatilis.

Lepidus[271]

Gen. et sp. nov.

Valid

Nesbitt & Ezcurra

Late Triassic

Dockum Group

 United States

A coelophysoid theropod. The type species is Lepidus praecisio.

Morelladon[272]

Gen. et sp. nov.

Valid

Gasulla et al.

Early Cretaceous (probably late Barremian)

Arcillas de Morella Formation

 Spain

An ornithopod dinosaur, a basal member of Styracosterna. The type species is Morelladon beltrani.

Mosaiceratops[273]

Gen. et sp. nov

Valid

Zheng, Jin & Xu

Late Cretaceous (early-middle Turonian—middle Campanian)

Xiaguan Formation

 China

A basal neoceratopsian. The type species is Mosaiceratops azumai.

Nebulasaurus[274]

Gen. et sp. nov

Valid

Xing et al.

Middle Jurassic (Aalenian or Bajocian)

Zhanghe Formation

 China

A eusauropod sauropod. The type species is Nebulasaurus taito.

Padillasaurus[275]

Gen. et sp. nov

Valid

Carballido et al.

Early Cretaceous (Barremian)

Paja Formation

 Colombia

A brachiosaurid sauropod. The type species is Padillasaurus leivaensis.

Probrachylophosaurus[276]

Gen. et sp. nov

Valid

Freedman Fowler & Horner

Late Cretaceous (Campanian)

Judith River Formation

 United States

A hadrosaurid. The type species is Probrachylophosaurus bergei.

Pulanesaura[277]

Gen. et sp. nov

Valid

McPhee et al.

Early Jurassic (probably late Hettangian to Sinemurian)

Upper Elliot Formation

 South Africa

A basal sauropod. The type species is Pulanesaura eocollum.

Qijianglong[278]

Gen. et sp. nov

Valid

Xing et al.

Late Jurassic

Suining Formation

 China

A mamenchisaurid sauropod. The type species is Qijianglong guokr.

Regaliceratops[279]

Gen. et sp. nov

Valid

Brown & Henderson

Late Cretaceous (probably middle Maastrichtian)

St. Mary River Formation

 Canada

A chasmosaurine ceratopsid. The type species is Regaliceratops peterhewsi.

Riabininohadros[280]

Gen. et comb. nov

Valid

Ulansky

Late Cretaceous (Maastrichtian)

Mt. Besh-Kosh

 Ukraine

A hadrosaurid of uncertain placement; new genus for "Orthomerus" weberi Riabinin, 1945.

Saurornitholestes sullivani[281]

Sp. nov

Valid

Jasinski

Late Cretaceous (late Campanian)

Kirtland Formation

 United States

A member of Dromaeosauridae, a species of Saurornitholestes.

Sefapanosaurus[282]

Gen. et sp. nov

Valid

Otero et al.

Late Triassic or Early Jurassic

Elliot Formation

 South Africa

A non-sauropod sauropodomorph. The type species is Sefapanosaurus zastronensis.

Sirindhorna[283]

Gen. et sp. nov

Valid

Shibata et al.

Early Cretaceous (Aptian)

Khok Kruat Formation

 Thailand

A basal member of Hadrosauroidea. The type species is Sirindhorna khoratensis.

Ugrunaaluk[284]

Gen. et sp. nov

Valid

Mori, Druckenmiller & Erickson

Late Cretaceous (early Maastrichtian)

Prince Creek Formation

 United States

A saurolophine hadrosaurid. The type species is Ugrunaaluk kuukpikensis.

Wendiceratops [285]

Gen. et sp. nov.

Valid

Evans & Ryan

Late Cretaceous (Campanian)

Oldman Formation

 Canada

A centrosaurine ceratopsid. The type species is Wendiceratops pinhornensis.

Yi[286]

Gen. et sp. nov

Valid

Xu et al.

Jurassic (Callovian or Oxfordian)

Tiaojishan Formation

 China

A scansoriopterygid theropod. The type species is Yi qi.

Zhenyuanlong[287]

Gen. et sp. nov.

Valid

LĂŒ & Brusatte

Early Cretaceous

Yixian Formation

 China

A dromaeosaurid theropod. The type species is Zhenyuanlong suni.

Birds

  • Balaur bondoc, originally interpreted as a member of Dromaeosauridae, is argued to be a basal member of Avialae by Cau, Brougham and Naish (2015).[288]
  • Estimates of likely mass and flight capability of the extinct Chatham duck (Anas chathamica), as indicated by measurements of major bones, are published by Williams (2015).[289]
  • New cranial material of the Oligo-Miocene relative of flamingos, Harrisonavis croizeti, is described by Torres et al. (2015).[290]
  • New specimens of the Eocene relative of swifts, Scaniacypselus szarskii, are described by Mayr (2015).[291]
  • A specimen of a stem-coliiform Masillacolius brevidactylus with preserved skull is described by Mayr (2015).[292]
  • A study of the phylogenetic relationships of Psittacopes, Pumiliornis and Morsoravis, indicating their close relationship to the passerines and extinct family Zygodactylidae, is published by Mayr (2015); the author also introduced a new name, Parapasseres, for the clade containg Passeriformes and Zygodactylidae but not Psittacopes.[293]
  • A study of preserved specimens of the Puerto Rican conure (previously considered to be an extinct subspecies of the Hispaniolan parakeet) is published by Olson (2015), who raises it to the rank of a separate species Psittacara maugei.[294]
  • A Dwarf Emu was found in the Miocene of Alcoota in Central Australia.[295]
  • Jones et al. (2015) describe a distal end of the left tibiotarsus of a member of the genus Caracara of uncertain specific assignment from the late Pleistocene of Argentina, estimated to be the largest member of the family Falconidae described thus far.[296]
  • A description of the skeletal anatomy of two well-preserved specimens of the dodo is published by Claessens, Meijer and Hume (2015).[297]
Name Novelty Status Authors Age Unit Location Notes Images

Aegypius tugarinovi [298]

Sp. nov.

Valid

Manegold & Zelenkov

Early Pliocene, Ruscinian

 Moldova

An Old World vulture, Accipitridae, related to the cinereous vulture.

Alopecoenas norfolkensis [299]

Sp. nov

Valid

Forshaw

Holocene

 Australia

A new scientific name for the Norfolk ground dove. The previous name used to refer to this species, Columba norfolciensis Latham (1801), was suppressed by ICZN in 2010.[300]

Anas kurochkini [301]

Sp. nov.

Valid

Zelenkov & Panteleyev

Late Miocene

 Russia

An Anatidae.

Archaeornithura [302]

Gen. et sp. nov.

Valid

Wang et al.

Hauterivian

Huajiying Formation

 China

A Hongshanornithidae. Type species A. meemannae.

Athene inexpectata [303]

Sp. nov.

Valid

Pavia, Manegold, & Haarhoff

Early Pliocene

Varswater Formation

 South Africa

A Strigidae

Circaetus haemusensis [304]

Sp. nov.

Valid

Boev

Early Pleistocene

 Bulgaria

A snake eagle

Cratoavis [305]

Gen. et sp. nov.

Valid

Carvalho et al.

Aptian

Crato Member, Santana Formation

 Brazil

A member of Euenantiornithes of uncertain phylogenetic placement.[305][306]
The type species is C. cearensis.

Dunhuangia [307]

Gen. et sp. nov.

Valid

Wang et al.

Early Cretaceous

Xiagou Formation

 China

A member of Enantiornithes. The type species is D. cuii.

Eopachypteryx [308]

Gen. et sp. nov.

Valid

Mayr

Eocene

Messel Formation

 Germany

A bird of uncertain phylogenetic placement.
Type species E. praeterita.

Feitianius [309]

Gen. et sp. nov

Valid

O’Connor et al.

Early Cretaceous (Aptian)

Xiagou Formation

 China

A member of Enantiornithes. The type species is Feitianius paradisi.

Foshanornis [310]

Gen. et sp. nov.

Valid

Zhao et al.

Early Eocene

Buxin Formation

 China

A bird of uncertain phylogenetic placement, possibly a relative of trogons. Type species F. songi.

Fumicollis [311]

Gen. et sp. nov.

Valid

Bell & Chiappe

Late Cretaceous (late Coniacian—early Campanian)

The Smoky Hill Member of the Niobrara Chalk

 United States

A member of Hesperornithiformes. The type species is Fumicollis hoffmani.

Gallinago azovica [301]

Sp. nov.

Valid

Zelenkov & Panteleyev

Middle Turolian

 Russia

A snipe, Scolopacidae.

Hakawai [312]

Gen. et sp. nov.

Valid[313]

De Pietri et al.

Early Miocene (19–16 Ma)

Bannockburn Formation

 New Zealand

A member of Scolopaci of uncertain phylogenetic placement. The type species is Hakawai melvillei.

Holbotia [314]

Gen. et sp. nov.

Valid[315]

Zelenkov & Averianov

Early Cretaceous

 Mongolia

A member of Enantiornithes, probably a relative of Gobipteryx. Type species H. ponomarenkoi.

Houornis [316]

Gen. et comb. nov.

Valid[317]

Wang & Liu

Early Cretaceous

Jiufotang Formation

 China

A member of Enantiornithes of uncertain phylogenetic placement; a new genus for "Cathayornis" caudatus Hou (1997).

Hypsipetes cowlesi [318]

Sp. nov.

Valid[319]

Hume

Holocene

 Rodrigues

A bulbul, Pycnonotidae

Juehuaornis [320]

Gen. et sp. nov.

Valid

Wang, Wang & Hu

Early Cretaceous

Jiufotang Formation

 China

A basal member of Ornithuromorpha. Type species J. zhangi.

Kuntur [321]

Gen. et sp. nov.

Valid

Stucchi et al.

Late Miocene

Pisco Formation

 Peru

A condor. Type species K. cardenasi.

Lavadytis [322]

Gen. et sp. nov.

Valid[323]

Stidham & Hilton

Miocene (16.1–14.6 Ma)

 United States

A member of Anatidae; a member or a relative of members of the subfamily Oxyurinae. The type species is Lavadytis pyrenae.

Llallawavis[324]

Gen. et sp. nov

Valid

Degrange et al.

Late Pliocene

Playa Los Lobos Allo Formation

 Argentina

A mesembriornithine phorusrhacid. Type species L. scagliai.

Makahala [325]

Gen. et sp. nov.

Valid

Mayr

Latest Eocene or early Oligocene

Makah Formation

 United States

A member of Procellariiformes. Type species M. mirae.

Mangystania[326]

Gen. et sp. nov

Valid

Zvonok, Zelenkov & Danilov

Eocene (Bartonian)

Shorym Formation

 Kazakhstan

Possibly a member of Suliformes. The type species is Mangystania humilicristata.

Miobaptus huzhiricus [327]

Sp. nov

Valid

Zelenkov

Miocene

 Russia

A grebe

Mioporphyrula [328]

Gen. et comb. nov

Valid

Zelenkov in Zelenkov & Kurochkin

Late Miocene

 Moldova

A member of Rallidae. A new genus for "Tertiariaporphyrula" lungi Kurochkin & Ganea (1972).

Mwalau [329]

Gen. et sp. nov.

Valid

Worthy et al.

Holocene

 Vanuatu

A megapode. The type species is M. walterlinii.

Namapsitta [330]

Gen. et sp. nov.

Valid

Mourer-Chauviré, Pickford, & Senut

Middle Eocene

 Namibia

A Psittaciformes incertae sedis. Type species N. praeruptorum.

Nyctisoma [331]

Gen. et sp. nov.

Valid

ElĆŒanowski & Zelenkov

Middle Miocene

Öoshin Formation

 Mongolia

An Ardeidae. Type species N. robusta.

Oligonomus [332]

Gen. et sp. nov.

Valid

De Pietri, Camens, & Worthy

Late Oligocene

 Australia

A member of Charadriiformes related to the plains-wanderer. Type species O. milleri.

Opisthodactylus horacioperezi [333]

Sp. nov.

Valid

Agnolin & Chafrat

Early Miocene

Chichinales Formation

 Argentina

A rheid ratite

Parapengornis [334]

Gen. et sp. nov.

Valid

Hu, O’Connor & Zhou

Early Cretaceous

Jiufotang Formation

 China

A member of Enantiornithes, a relative of Pengornis. Type species P. eurycaudatus.

Patagorhacos [333]

Gen. et sp. nov.

Valid

Agnolin & Chafrat

Early Miocene

Chichinales Formation

 Argentina

A member of Phorusrhacidae. Type species P. terrificus.

Petrosushkinia [328]

Nom. nov

Valid

Zelenkov in Zelenkov & Kurochkin

Late Miocene

 Kazakhstan

A member of Falconidae. A replacement name for Sushkinia Tugarinov (1935) (preoccupied).

Porzana botunensis [335]

Sp. nov.

Valid

Boev

Early Pleistocene

 Bulgaria

A member of Rallidae, a species of crake.

Pterodroma imberi [336]

Sp. nov.

Valid

Tennyson, Cooper & Shepherd

Holocene

 New Zealand

A member of Procellariidae, a species of gadfly petrel.

Pterygornis [337]

Gen. et sp. nov.

Valid [338]

Wang, Hu & Li

Early Cretaceous

Jiufotang Formation

 China

A member of Enantiornithes. Type species P. dapingfangensis.

Rallus adolfocaesaris [339]

Sp. nov.

Valid

Alcover et al.

Holocene

 Madeira (Porto Santo Island)

A member of Rallidae, a species of Rallus

Rallus carvaoensis [339]

Sp. nov.

Valid

Alcover et al.

Holocene

 Azores (SĂŁo Miguel Island)

A member of Rallidae, a species of Rallus

Rallus lowei [339]

Sp. nov.

Valid

Alcover et al.

Holocene

 Madeira (Madeira Island)

A member of Rallidae, a species of Rallus

Rallus minutus [339]

Sp. nov.

Junior homonym

Alcover et al.

Holocene

 Azores (SĂŁo Jorge Island)

A member of Rallidae, a species of Rallus. The specific name is a junior homonym of Rallus minutus Pallas (1776), Rallus minutus Gmelin (1789) and Rallus minutus Forster (1844). Alcover et al. (2016) coined a replacement name Rallus nanus.[340]

Rallus montivagorum [339]

Sp. nov.

Valid

Alcover et al.

Holocene

 Azores (Pico Island)

A member of Rallidae, a species of Rallus

Ramphastosula aguirrei [341]

Sp. nov.

Valid

Stucchi, Varas-Malca & Urbina-Schmitt

Miocene

Pisco Formation

 Peru

A member of Sulidae, a species of Ramphastosula.

Scolopax brachycarpa [342]

Sp. nov.

Valid

Takano & Steadman

Late Holocene

 Haiti

A woodcock.

Scopelortyx [330]

Gen. nov. et Sp. nov.

Valid

Mourer-Chauviré, Pickford, & Senut

Middle Eocene

 Namibia

A Paraortygidae, Galliformes. Type species S. klinghardtensis.

Shiriyanetta [343]

Gen. et sp. nov.

Valid

Watanabe & Matsuoka

Pleistocene

 Japan

A duck. The type species is Shiriyanetta hasegawai.

Spheniscus anglicus [344]

Sp. nov.

Valid

Benson

Late Miocene

BahĂ­a Inglesa Formation

 Chile

A banded penguin.

Stemec [345]

Gen. et sp. nov.

Valid

Kaiser, Watanabe & Johns

Late Oligocene

Sooke Formation

 Canada

A member of Plotopteridae. The type species is Stemec suntokum.

Sula brandi [341]

Sp. nov.

Valid

Stucchi, Varas-Malca & Urbina-Schmitt

Miocene

Pisco Formation

 Peru

A booby.

Sula figueroae [341]

Sp. nov.

Valid

Stucchi, Varas-Malca & Urbina-Schmitt

Miocene

Pisco Formation

 Peru

A booby.

Tyto cravesae [346]

Sp. nov.

Valid

SuĂĄrez & Olson

Quaternary (probably late Pleistocene)

 Cuba

A barn-owl, a species of Tyto.

Tyto richae [303]

Sp. nov

Valid

Pavia, Manegold, & Haarhoff

Early Pliocene

Varswater Formation

 South Africa

A Tytonidae barn-owl.

Yuanjiawaornis [347]

Gen. et sp. nov.

Valid

Hu et al.

Early Cretaceous

Jiufotang Formation

 China

A member of Enantiornithes. Type species Yuanjiawaornis viriosus.

Pterosaurs

  • A study on the terrestrial locomotion of non-pterodactyloid pterosaurs is published by Witton (2015).[348]
  • A specimen of Rhamphorhynchus muensteri with preserved soft tissues or impressions of soft tissues, stomach contents and possibly a coprolite is described by Hone et al. (2015).[349]
Name Novelty Status Authors Age Unit Location Notes Images

Arcticodactylus[350]

Gen. et comb. nov

Valid

Kellner

Late Triassic (?Norian-Rhaetian)

Fleming Fjord Formation

 Greenland

A non-pterodactyloid pterosaur of uncertain phylogenetic placement; a new genus for "Eudimorphodon" cromptonellus Jenkins et al. (2001).

Austriadraco[350]

Gen. et sp. nov

Valid

Kellner

Late Triassic (late Norian)

Seefeld Formation

 Austria

A non-pterodactyloid pterosaur of uncertain phylogenetic placement. The type species is Austriadraco dallavecchiai.

Banguela[351]

Gen. et sp. nov

Disputed

Headden & Campos

Early Cretaceous

Romualdo Formation

 Brazil

The type species is Banguela oberlii. Originally described as a dsungaripterid; PĂȘgas, Costa & Kellner (2018) considered the genus Banguela to be a junior synonym of the genus Thalassodromeus, and transferred the species B. oberlii to the latter genus.[352]

Bergamodactylus[350]

Gen. et sp. nov

Disputed

Kellner

Late Triassic (Alaunian)

Calcari di Zorzino Formation

 Italy

Originally interpreted as a relative of Campylognathoides. The type species is Bergamodactylus wildi. Dalla Vecchia (2018) considers B. wildi to be a junior synonym of Carniadactylus rosenfeldi.[353]

Cimoliopterus dunni[354]

Sp. nov

Valid

Myers

Late Cretaceous (late Cenomanian)

Britton Formation

 United States

A member of Pterodactyloidea, probably a pteranodontoid; a species of Cimoliopterus.

Daohugoupterus[355]

Gen. et sp. nov

Valid

Cheng et al.

Late Jurassic

Daohugou Beds

 China

A non-pterodactyloid pterosaur. The type species is Daohugoupterus delicatus.

Linlongopterus[356]

Gen. et sp. nov

Valid

Rodrigues et al.

Early Cretaceous (Aptian)

Jiufotang Formation

 China

A member of Pteranodontoidea, possibly related to anhanguerians. The type species is Linlongopterus jennyae.

Orientognathus[357]

Gen. et sp. nov

Valid

LĂŒ et al.

Late Jurassic

Tuchengzi Formation

 China

A rhamphorhynchid. The type species is Orientognathus chaoyngensis.

Rhamphorhynchus etchesi[358]

Sp. nov

Valid

O'Sullivan & Martill

Late Jurassic

Kimmeridge Clay Formation

 United Kingdom

A species of Rhamphorhynchus.

Other

Other reptiles

Name Novelty Status Authors Age Unit Location Notes Images

Eohyosaurus[360]

Gen. et sp. nov

Valid

Butler et al.

Middle Triassic (early Anisian)

Burgersdorp Formation

 South Africa

A basal rhynchosaur. The type species is Eohyosaurus wolvaardti.

Erpetonyx[361]

Gen. et sp. nov

Valid

Modesto et al.

Carboniferous (Gzhelian)

Egmont Bay Formation

 Canada

A relative of bolosaurids. The type species is Erpetonyx arsenaultorum.

Glaurung[362]

Gen. et sp. nov

Valid

Bulanov & Sennikov

Late Permian

 Germany

A member of Weigeltisauridae. The type species is Glaurung schneideri.

Opisthodontosaurus[363]

Gen. et sp. nov

Valid

Reisz et al.

Early Permian

Garber Formation

 United States

A member of Captorhinidae. The type species is Opisthodontosaurus carrolli.

Pappochelys[364]

Gen. et sp. nov

Valid

Schoch & Sues

Middle Triassic (Ladinian)

Erfurt Formation

 Germany

A stem-turtle. The type species is Pappochelys rosinae.

"Thalassodromeus" sebesensis[365]

Sp. nov

Valid

Grellet-Tinner & Codrea

Late Cretaceous (Maastrichtian)

 Romania

A reptile of uncertain phylogenetic placement; initially classified as a pterosaur and a species of Thalassodromeus,[365] but subsequently argued to be a turtle and a junior synonym of Kallokibotion bajazidi.[366]

Synapsids

Non-mammalian synapsids

  • Redescription of the anatomy of Tiarajudens eccentricus is published by Cisneros et al. (2015), who compare the species with Anomocephalus africanus.[367]
Name Novelty Status Authors Age Unit Location Notes Images

Endothiodon tolani[368]

Sp. nov

Valid

Cox & Angielczyk

Permian

Ruhuhu Formation

 Tanzania

A dicynodont, a species of Endothiodon.

Ichibengops[369]

Gen. et sp. nov

Valid

Huttenlocker, Sidor & Angielczyk

Late Permian

Madumabisa Mudstone Formation

 Zambia

A therocephalian. The type species is Ichibengops munyamadziensis.

Lende[370]

Gen. et sp. nov

Valid

Kruger et al.

Late Permian

Chiweta Beds

 Malawi

A burnetiamorph biarmosuchian therapsid. The type species is Lende chiweta.

Nothogomphodon sanjiaoensis[371]

Sp. nov

Valid

Liu & Abdala

Middle Triassic

Ermaying Formation

 China

A therocephalian, a species of Nothogomphodon.

Ruberodon[372]

Gen. et sp. nov

Valid

Ray

Late Triassic

Tiki Formation

 India

A traversodontid cynodont. The type species is Ruberodon roychowdhurii.

Sinokannemeyeria baidaoyuensis[373]

Sp. nov

Valid

Liu

Middle Triassic

Tongchuan Formation

 China

A kannemeyeriiform dicynodont, a species of Sinokannemeyeria.

Mammals

  • Description of exceptionally preserved specimens of the pantodont species Alcidedorbignya inopinata and a study of its phylogenetic relationships is published by De Muizon et al. (2015).[374]
  • A study on the phylogenetic relationships of the family Nyctitheriidae based on new fossil remains of Plagioctenodon rosei and Plagioctenodon thewisseni is published by Manz et al. (2015).[375]
  • Studies of the phylogenetic relationships of litopterns and notoungulates within Placentalia, indicating that their closest living relatives are odd-toed ungulates, are published by Welker et al. (2015)[376] and Buckley (2015).[377]
  • A study of the phylogenetic relationships of the Pleistocene camelid genus Camelops, based on genomic data extracted from its bones, is published by Heintzman et al. (2015).[378]
  • A study on the feeding habits of extant and fossil members of the family Canidae belonging to the tribe Canini, as indicated by skull size and shape, is published by Meloro, Hudson & Hook (2015).[379]
  • A study on the population dynamics of the cave lion (Panthera spelaea) during the Late Pleistocene is published by Ersmark et al. (2015).[380]
  • Description of tarsals attributed to Purgatorius and a study on the phylogenetic relationships of this mammal is published by Chester et al. (2015).[381]
  • A study on the age of the holotype specimen of Darwinius masillae at the time of death and on the life history of the species is published by LĂłpez-Torres, Schillaci & Silcox (2015).[382]
  • Cranial endocast of Victoriapithecus macinessi is reconstructed by Gonzales et al. (2015).[383]
  • A study evaluating the evidence of association between extinctions or replacements of large mammals in the Northern Hemisphere in the Late Pleistocene with warming events is published by Cooper et al. (2015).[384]
Name Novelty Status Authors Age Unit Location Notes Images

Antawallathentes[385]

Gen. et 2 sp. nov

Valid

RincĂłn et al.

Oligocene (Deseadan)

Salla Formation

 Bolivia

A palaeothentid marsupial (a relative of the shrew opossums). Genus contains two species: Antawallathentes illimani and Antawallathentes quimsacruza.

Archaeonothos[386]

Gen. et sp. nov

Valid

Beck

Early Eocene

Murgon fossil site

 Australia

A metatherian of uncertain phylogenetic placement. The type species is Archaeonothos henkgodthelpi.

Ganguroo robustiter[387]

Sp. nov

Valid

Cooke et al.

Miocene

Riversleigh World Heritage Area

 Australia

A basal member of Macropodidae, a species of Ganguroo.

Gaylordia mater[388]

Sp. nov

Valid

De Oliveira & Goin

Itaboraian

ItaboraĂ­ Formation

 Brazil

A metatherian of uncertain phylogenetic placement, possibly a relative of Pucadelphys; a species of Gaylordia.

Lotheridium[389]

Gen. et sp. nov

Valid

Bi et al.

Late Cretaceous

Qiupa Formation

 China

A member of Deltatheridiidae. The type species is Lotheridium mengi.

Lycopsis padillai[390]

Sp. nov

Valid[391]

Suarez et al.

Miocene

Castilletes Formation

 Colombia

A member of Sparassodonta, a species of Lycopsis.

Madju[392]

Gen. et 2 sp. nov

Valid

Travouillon et al.

Late Oligocene to early late Miocene

 Australia

A bandicoot. Genus contains two species: Madju variae and Madju encorensis.

Nimbavombatus[393]

Gen. et sp. nov

Valid

Brewer et al.

Early Miocene

Riversleigh World Heritage Area

 Australia

A wombat. The type species is Nimbavombatus boodjamullensis.

Oklatheridium wiblei[394]

Sp. nov

Valid

Cifelli & Davis

Early Cretaceous (Albian)

Cloverly Formation

 United States

A member of Deltatheroida, a species of Oklatheridium.

Sedophascolomys[395]

Gen. et comb. nov

Valid

Louys

Pliocene to Pleistocene

 Australia

A wombat; a new genus for "Phascolomys" medius Owen (1872).

Sparassocynus maimarai[396]

Sp. nov

Valid

Abello et al.

Late Miocene

MaimarĂĄ Formation

 Argentina

A sparassocynid didelphoid (a relative of the opossums), a species of Sparassocynus.

Tsagandelta[397]

Gen. et sp. nov

Valid

Rougier, Davis & Novacek

Late Cretaceous

Baynshiree Formation

 Mongolia

A member of Deltatheroida. The type species is Tsagandelta dashzevegi.

Wabularoo prideauxi[398]

Sp. nov

Valid

Travouillon, Archer & Hand

Early Miocene

Riversleigh World Heritage Area

 Australia

A member of Macropodoidea, a species of Wabularoo.

Name Novelty Status Authors Age Unit Location Notes Images

Agilodocodon[399]

Gen. et sp. nov

Valid

Meng et al.

Middle Jurassic

"Daohugou site in Tiaojishan Formation (...), also known as Jiulongshan Formation"[399]

 China

A docodont. The type species is Agilodocodon scansorius.

Argaliatherium[394]

Gen. et sp. nov

Valid

Cifelli & Davis

Early Cretaceous (Albian)

Cloverly Formation

 United States

A basal member of Tribosphenida. The type species is Argaliatherium robustum.

Carinalestes[394]

Gen. et sp. nov

Valid

Cifelli & Davis

Early Cretaceous (Albian)

Cloverly Formation

 United States

A basal member of Tribosphenida. The type species is Carinalestes murensis.

Docodon apoxys[400]

Sp. nov

Valid

Rougier et al.

Late Jurassic

Morrison Formation

 United States

A docodont, a species of Docodon.

Docofossor[401]

Gen. et sp. nov

Valid

Luo et al.

Late Jurassic

Tiaojishan Formation

 China

A docodont. The type species is Docofossor brachydactylus.

Gobiconodon haizhouensis[402]

Sp. nov

Valid

Kusuhashi et al.

Early Cretaceous

Fuxin Formation

 China

A gobiconodontid, a species of Gobiconodon.

Gobiconodon tomidai[402]

Sp. nov

Valid

Kusuhashi et al.

Early Cretaceous

Shahai Formation

 China

A gobiconodontid, a species of Gobiconodon.

Kimbetopsalis[403]

Gen. et sp. nov

Valid

Williamson et al.

Early Paleocene (Puercan)

Nacimiento Formation

 United States

A taeniolabidid multituberculate. The type species is Kimbetopsalis simmonsae.

Spinolestes[404]

Gen. et sp. nov

Valid

Martin et al.

Early Cretaceous (latest Barremian)

Calizas de La Huergina Formation

 Spain

A gobiconodontid. The type species is Spinolestes xenarthrosus.

Tirotherium[405]

Gen. et sp. nov

Valid

Montellano-Ballesteros & Fox

Late Cretaceous (late Santonian to early Campanian)

Milk River Formation

 Canada

An early member of Boreosphenida, probably a member of Picopsidae. The type species is Tirotherium aptum.

Valenopsalis[403]

Gen. et comb. nov

Valid

Williamson et al.

Early Paleocene (Puercan)

Bug Creek Anthills and Bug Creek West

 United States

A taeniolabidoid multituberculate; a new genus for "Catopsalis" joyneri Sloan & Van Valen (1965).

Yubaatar[406]

Gen. et sp. nov

Valid

Xu et al.

Late Cretaceous

Qiupa Formation

 China

A cimolodont multituberculate. The type species is Yubaatar zhongyuanensis.

Name Novelty Status Authors Age Unit Location Notes Images

Eionaletherium[407]

Gen. et sp. nov.

Valid

RincĂłn et al.

Late Miocene

Urumaco Formation

 Venezuela

A mylodontoid sloth. The type species is Eionaletherium tanycnemius.

Tonnicinctus[408]

Gen. et sp. nov.

Valid

GĂłis et al.

Pleistocene (Ensenadan to Lujanian)

 Argentina

A member of Pampatheriidae. The type species is Tonnicinctus mirus.

Name Novelty Status Authors Age Unit Location Notes Images

Arenagale[409]

Gen. et sp. nov

Valid

Pickford

Eocene (Bartonian)

Eocliff Limestone

 Namibia

A tenrec. The type species is A. calcareus.

Callistosiren[410]

Gen. et sp. nov.

Valid

VĂ©lez-Juarbe & Domning

Late Oligocene

Lares Limestone

 United States ( Puerto Rico)

A dugong. The type species is Callistosiren boriquensis.

Diamantochloris[411]

Gen. et sp. nov

Valid

Pickford

Eocene (Lutetian)

Black Crow Limestone

 Namibia

A golden mole. The type species is D. inconcessus.

Namachloris[412]

Gen. et sp. nov

Valid

Pickford

Eocene (Bartonian)

Eocliff Limestone

 Namibia

A golden mole. The type species is N. arenatans.

Namagale[409]

Gen. et sp. nov

Valid

Pickford

Eocene (Bartonian)

Eocliff Limestone

 Namibia

A member of Potamogalidae. The type species is N. grandis.

Palaeoloxodon xylophagou[413]

Sp. nov

Valid

Athanassiou et al.

Late middle Pleistocene

 Cyprus

A member of Elephantidae, a species of Palaeoloxodon.

Protanancus brevirostris[414]

Sp. nov.

Valid

Wang et al.

Early Miocene

Linxia Basin

 China

A gomphothere, a species of Protanancus.

Rupestrohyrax[415]

Gen. et sp. nov

Valid

Pickford

Eocene (Bartonian)

Eoridge Limestone

 Namibia

A hyrax related to Titanohyrax. The type species is R. palustris.

Sperrgale[409]

Gen. et sp. nov

Valid

Pickford

Eocene (Bartonian)

Eocliff Limestone

 Namibia

A tenrec. The type species is S. minutus.

Thyrohyrax libycus[416]

Sp. nov

Valid

Coster et al.

Early Oligocene

 Libya

A saghatheriid hyracoid, a species of Thyrohyrax.

Name Novelty Status Authors Age Unit Location Notes Images

Cardioderma leakeyi[417]

Sp. nov

Valid

Gunnell et al.

Early Pleistocene

Olduvai Gorge locality

 Tanzania

A member of Megadermatidae related to the heart-nosed bat.

Marnenycteris[418]

Gen. et sp. nov

Valid

Hand et al.

Eocene (Ypresian)

 France

A bat related to Onychonycteris. The type species is Marnenycteris michauxi.

Mystacina miocenalis[419]

Sp. nov.

Valid

Hand, Lee, Worthy & Archer in Hand et al.

Early Miocene (Altonian, 19–16 Ma)

 New Zealand

A New Zealand short-tailed bat.

Myzopoda africana[417]

Sp. nov

Valid

Gunnell et al.

Early Pleistocene

Olduvai Gorge locality

 Tanzania

A species of Myzopoda.

Nycticeinops serengetiensis[417]

Sp. nov

Valid

Gunnell et al.

Early Pleistocene

Olduvai Gorge locality

 Tanzania

A vesper bat related to the Schlieffen's bat.

Otonycteris rummeli[420]

Sp. nov

Valid

Rosina

Late Miocene

 Ukraine

A vesper bat related to the desert long-eared bat.

Scotoecus olduvensis[417]

Sp. nov

Valid

Gunnell et al.

Early Pleistocene

Olduvai Gorge locality

 Tanzania

A vesper bat, a species of Scotoecus.

Name Novelty Status Authors Age Unit Location Notes Images

Boreohippidion[421]

Gen. et comb. nov.

Valid

Avilla, Bernardes & Mothé

Early Pliocene (late Hemphillian)

 United States

A member of Equidae; a new genus for "Onohippidium" galushai Macfadden & Skinner (1979).

Cambaylophus[422]

Gen. et sp. nov.

Valid

Kapur & Bajpai

Early Eocene

Cambay Shale

 India

A member of Tapiroidea of uncertain phylogenetic placement. The type species is C. vastanensis.

Chasmotherium depereti[423]

Sp. nov

Valid

Remy

Eocene (Bartonian)

 France

A relative of Hyrachyus, a species of Chasmotherium.

Eolophiodon[424]

Gen. et sp. nov.

Valid

Robinet et al.

Early Eocene

 France

A relative of Lophiodon. The type species is Eolophiodon laboriense.

Pachynolophus gaytei[423]

Sp. nov

Valid

Remy

Eocene (Bartonian)

 France

A member of Palaeotheriidae, a species of Pachynolophus.

Persiatherium[425]

Gen. et sp. et comb. nov

Valid

Pandolfi

Late Miocene

 China
 Iran

A rhinoceros related to Aceratherium. The type species is P. rodleri; genus also includes"Aceratherium" huadeensis Qiu (1979).

Plesiohipparion shanxiense[426]

Sp. nov.

Valid

Bernor, Sun & Chen

Early Pleistocene

 China

A member of Equidae related to Hipparion; a species of Plesiohipparion.

Vastanolophus[427]

Gen. et sp. nov

Valid

Smith et al.

Eocene (Ypresian)

Cambay Shale Formation

 India

A member of Tapiroidea, possibly a member of Helaletidae. The type species is Vastanolophus holbrooki.

Name Novelty Status Authors Age Unit Location Notes Images

Anthracokeryx naduongensis[428]

Sp. nov

Valid

Ducrocq et al.

Eocene

Na Duong Formation
Youganwo Formation[429]

 China[429]
 Vietnam

An anthracothere, a species of Anthracokeryx.

Bothriogenys langsonensis[428]

Sp. nov

Valid

Ducrocq et al.

Eocene

Na Duong Formation

 Vietnam

An anthracothere, a species of Bothriogenys.

Bothriogenys vietnamensis[428]

Sp. nov

Valid

Ducrocq et al.

Eocene

Na Duong Formation

 Vietnam

An anthracothere, a species of Bothriogenys.

Eotragus lampangensis[430]

Sp. nov

Valid

Suraprasit et al.

Late Middle Miocene (13.4-13.2 Ma)

Nakhaem Formation

 Thailand

A bovid, a species of Eotragus.

Epirigenys[431]

Gen. et sp. nov

Valid

Lihoreau et al.

Oligocene

Lokone Sandstone Formation

 Kenya

A relative of hippopotamuses. The type species is Epirigenys lokonensis.

Eucladoceros montenegrensis[432]

Sp. nov

Valid

Van der Made & Dimitrijević

Early Pleistocene

 Montenegro

A deer, a species of Eucladoceros.

Euprox grandis[433]

Sp. nov

Valid

Hou

Late Miocene

Liushu Formation